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Neuroscience, PhD

School of Medicine

Catalogue Home

  • Explore our Programs
  • University-​wide Policies and Information
    • Academic Policies and Information
      • Academic Calendar
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        • Financial Aid
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    • Office of Institutional Equity
      • Discrimination and Harassment Policy and Procedures
      • Equal Opportunity and Title IX Notice
      • Sexual Misconduct Policy and Procedures
    • Rights, Privileges, and Responsibilities
      • Academic Grievance Policy: Students and Postdoctoral Fellows
      • New Child Accommodations for Full-​Time Graduate Students and Postdoctoral Trainees
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  • Bloomberg School of Public Health
    • Academic Calendar
    • Admission
    • CEPH Requirements
    • Departments
      • Department of Biochemistry and Molecular Biology
        • Biochemistry and Molecular Biology, MHS
        • Biochemistry and Molecular Biology, ScM
        • Biochemistry and Molecular Biology, PhD
        • Non-​Degree Training
      • Department of Biostatistics
        • Biostatistics, MHS
        • Biostatistics, ScM
        • Biostatistics, PhD
      • Department of Environmental Health and Engineering
        • Environmental Health, MHS
        • Environmental Health, SCM
        • Toxicology for Human Risk Assessment, MS
        • Environmental Health, PhD
        • Non-​Degree Training
      • Department of Epidemiology
        • Epidemiology, MHS
        • Epidemiology, ScM
        • Epidemiology, PhD
        • Non-​Degree Training
      • Department of Health, Behavior and Society
        • Health Education and Health Communication, MSPH
        • Genetic Counseling, ScM
        • Health, Behavior, and Society, MHS
        • Social and Behavioral Sciences, PhD
        • Non-​Degree Training
      • Department of Health Policy and Management
        • Health Administration, MHA
        • Health Economics and Outcomes Research, MHS
        • Health Finance and Management, MHS
        • Health Policy, MSPH
        • Health Policy and Management, PhD
        • Health Policy and Management, DrPH (Tsinghua)
        • Non-​Degree Training
      • Department of International Health
        • Global Health Economics, MHS
        • International Health, MSPH
        • International Health, MSPH, Human Nutrition-​Dietitian
        • International Health, MA/​MSPH
        • International Health, PhD
        • Non-​Degree Training
      • Department of Mental Health
        • Mental Health, MHS
        • Mental Health, PhD
        • Non-​Degree Training
      • Department of Molecular Microbiology &​ Immunology
        • Molecular Microbiology &​ Immunology, MHS
        • Molecular Microbiology &​ Immunology, ScM
        • Molecular Microbiology &​ Immunology, PhD
        • Non-​Degree Training
      • Department of Population, Family and Reproductive Health
        • Population, Family and Reproductive Health, MHS
        • Population, Family and Reproductive Health, MHS Online
        • Population, Family and Reproductive Health, MSPH
        • Population, Family and Reproductive Health, PhD
      • Doctor of Public Health (DrPH)
      • Graduate Training Programs in Clinical Investigation
        • Clinical Investigation, MHS
        • Clinical Investigation, PhD
        • Clinical Investigation, ScM
      • Master of Arts in Public Health Biology
      • Master of Bioethics
      • Master of Public Health Program
        • DNP/​MPH
        • DVM/​MPH
        • JD/​MPH
        • LLM/​MPH
        • MBA/​MPH with China Europe International Business School
        • MD/​MPH
        • MPH/​MBA
        • MSW/​MPH
      • MAS-​Office
        • Master of Applied Science in Patient Safety and Healthcare Quality
        • Master of Applied Science in Population Health Management
        • Master of Applied Science in Spatial Analysis for Public Health
      • Bachelor's/​Master's Degrees
      • MD/​PhD
      • PhD/​MBA
      • Residency Programs
        • General Preventive Medicine Residency Program
        • Occupational and Environmental Medicine Residency
    • Certificates
      • Adolescent Health, Certificate
      • Bioethics, Certificate
      • Climate and Health, Certificate
      • Clinical Trials, Certificate
      • Community-​Based Public Health, Certificate
      • Demographic Methods, Certificate
      • Environmental and Occupational Health, Certificate
      • Epidemiology for Public Health Professionals, Certificate
      • Evaluation: International Health Programs, Certificate
      • Food Systems, the Environment &​ Public Health, Certificate
      • Gender and Health, Certificate
      • Gerontology, Certificate
      • Global Digital Health, Certificate
      • Global Health, Certificate
      • Health Communication, Certificate
      • Health Disparities and Health Inequality, Certificate
      • Health Education, Certificate
      • Health Finance and Management, Certificate
      • Healthcare Epidemiology and Infection Prevention and Control, Certificate
      • Humane Sciences and Toxicology Policy, Certificate
      • Humanitarian Health, Certificate
      • Implementation Science and Research Practice, Certificate
      • Indigenous Public Health Certificate
      • Infectious Disease Dynamics, Analytics, and Modeling Certificate
      • Injury and Violence Prevention, Certificate
      • Leadership for Public Health and Healthcare, Certificate
      • Lesbian, Gay, Bisexual, Transgender, and Queer (LGBTQ) Public Health, Certificate
      • Maternal and Child Health, Certificate
      • Mental Health Policy, Economics and Services, Certificate
      • Pharmacoepidemiology and Drug Safety, Certificate
      • Population and Health, Certificate
      • Population Health Management, Certificate
      • Product Stewardship for Sustainability, Certificate
      • Public Health Advocacy, Certificate
      • Public Health Economics, Certificate
      • Public Health Informatics, Certificate
      • Public Health Preparedness, Certificate
      • Public Health, Human Rights, and Law, Certificate
      • Public Mental Health Research, Certificate
      • Quality, Patient Safety, and Outcomes Research, Certificate
      • Rigor, Reproducibility and Responsibility in Scientific Practice, Certificate
      • Risk Sciences and Public Policy, Certificate
      • Social Epidemiology, Certificate
      • Spatial Analysis for Public Health, Certificate
      • Training Certificate in Public Health
      • Tropical Medicine, Certificate
      • Vaccine Science and Policy, Certificate
    • Policies
      • Academic
        • Academic Ethics Code
        • Compliance Line
        • Grade Appeal Policy
        • Grading System
        • Graduation Policy
        • Interdivisional Registration
        • Multi-​Term Course Policy
        • Post-​Doctoral Fellow Student Status
        • Student Grievance Policy
        • Voluntary Leave of Absence Policy
      • Research
        • Animal Research
        • Human Subjects Research
        • Worker's Compensation
  • Carey Business School
    • Admission
      • Master’s Programs
      • Certificate Programs
      • International Student Admission Policy
      • Verification of Credentials
      • Other Admission Policies
    • Degrees and Certificates
      • Artificial Intelligence for Business, Graduate Certificate
      • Business Administration (Accelerated), MBA
      • Business Administration (Executive), MBA
      • Business Administration (Flexible), MBA
      • Business Administration (Full Time), MBA
      • Business Analytics and Artificial Intelligence, Master of Science
      • Business Analytics and Artificial Intelligence (Part Time), Master of Science
      • Business Analytics and Risk Management, Graduate Certificate
      • Design Leadership, MBA/​MA Dual Degree
      • Digital Marketing, Graduate Certificate
      • Entrepreneurial Marketing, Graduate Certificate
      • Finance, Master of Science
      • Finance, Master of Science, Financial Econometrics Concentration
      • Finance (Part Time), Master of Science
      • Financial Management, Graduate Certificate
      • Financial Management, Graduate Certificate, Investments, Graduate Certificate, Applied Economics, MS
      • Health Care Management (Part Time), Master of Science
      • Health Care Management, Master of Science
      • Healthcare Management, Innovation, and Technology, Graduate Certificate
      • Information Systems and Artificial Intelligence for Business, Master of Science
      • Information Systems and Artificial Intelligence for Business (Part Time), Master of Science
      • Investments, Graduate Certificate
      • Management, Master of Science
      • Management (Part Time), Master of Science
      • Marketing, Master of Science
      • Marketing, Master of Science, Marketing Analytics Concentration
      • Marketing (Part Time), Master of Science
      • MBA/​Applied Economics, MS Dual Degree
      • MBA/​Biotechnology, MS Dual Degree
      • MBA/​Communication, MA Dual Degree
      • MBA/​DNP Dual Degree
      • MBA/​Government, MA Dual Degree
      • MBA/​Healthcare Organizational Leadership, MSN Dual Degree
      • MBA/​Health Care Management, MS Dual Degree
      • MBA/​JD Dual Degree
      • MBA/​MA in International Relations
      • MBA/​MD Dual Degree
      • MBA/​MPH Dual Degree
      • MBA/​PharmD Dual Degree
      • PhD/​MBA Dual Degree
      • Real Estate and Infrastructure (Part Time), Master of Science
      • Real Estate and Infrastructure, Master of Science
      • Business, Minor
    • Policies and Resources
      • Academic Calendar
      • Academic Ethics Policy
      • Academic Progress and Standards
      • Changing Degree Program
      • Grading Policy
      • Graduation
      • Attendance Policy
      • Leave of Absence
      • Registration
      • Student Accounts
      • Transfer of Graduate Credit
      • Waiver Exams
  • Peabody Institute
    • General Information, Procedures and Regulations
      • Introduction and Nomenclature
      • Mission
      • Accreditation
      • Links
      • Honor Societies
    • Procedural Information
      • Applicability
      • Studio Assignments
      • Course Numbering
      • Large Ensemble Participation
      • Competitions
      • Recitals
      • Academic Advising
      • Inter-​Institutional Academic Arrangements
      • Study Abroad Program
      • Outside Instruction and Public Performance
    • Academic Regulations
      • Applicability
      • Academic Code of Conduct
      • Program Classification, Status, and Credit Limits
      • Sources of Credit
      • Grading System and Regulations
      • Dean's List Criteria
      • Academic Standing
      • Registration Regulations
      • Attendance and Absences
      • Interruption of Degree Work
      • Graduation Eligibility
    • Degree and Diploma Programs
      • Bachelor of Fine Arts in Dance (BFA)
      • Bachelor of Music (BM)
        • Curricula
          • Bachelor of Music in Composition
          • Bachelor of Music in Hip Hop
          • Bachelor of Music in Jazz Performance
          • Bachelor of Music in Music Education
          • Bachelor of Music in Music for New Media
          • Bachelor of Music in Performance
            • Bachelor of Music in Performance
            • Bachelor of Music in Performance -​ Computer Music
            • Bachelor of Music in Performance -​ Guitar
            • Bachelor of Music in Performance -​ Harpsichord
            • Bachelor of Music in Performance -​ Historical Performance
            • Bachelor of Music in Performance -​ Orchestral Instruments
            • Bachelor of Music in Performance -​ Organ
            • Bachelor of Music in Performance -​ Piano
            • Bachelor of Music in Performance -​ Voice
          • Bachelor of Music in Recording Arts &​ Sciences
        • Minors
          • Business of Music, Minor
          • Directed Studies, Minor
          • Historical Performance, Minor
          • Historical Performance: Voice, Minor
          • Liberal Arts, Minor
          • Minors Offered at Other JHU Schools
          • Music Theory, Minor
          • Musicology, Minor
        • Combined Degree Programs
          • Peabody-​Homewood Double Degree Program
        • Accelerated Graduate Degrees
          • Five-​Year BM/​MM Program
          • Five-​Year BMRA/​MA Program
            • Five-​Year BM/​MA: Music for New Media Variant
      • Master of Music (MM)
        • Master of Music, Composition
        • Master of Music, Electronics and Computer Music
        • Master of Music, Film and Game Scoring
        • Master of Music: Performance
          • Master of Music, Performance -​ Choral Conducting specialization
          • Master of Music, Performance -​ Guitar specialization
          • Master of Music, Performance -​ Harpsichord specialization
          • Master of Music, Performance -​ Historical Performance Instruments specialization
          • Master of Music, Performance -​ Historical Performance Voice specialization
          • Master of Music, Performance -​ Jazz specialization
          • Master of Music, Performance -​ Orchestral Conducting specialization
          • Master of Music, Performance -​ Orchestral Instruments specialization
          • Master of Music, Performance -​ Organ specialization
          • Master of Music, Performance -​ Piano specialization
          • Master of Music, Performance -​ Wind Conducting specialization
          • Master of Music, Performance -​ Voice specialization
        • Master of Music: Academic Majors
          • Performance, Master of Music -​ Pedagogy emphasis
          • Music Education, Master of Music
          • Musicology, Master of Music
          • Music Theory Pedagogy, Master of Music
        • Master of Music: Low Residency
      • Master of Arts (MA)
        • Audio Sciences: Acoustics, Master of Arts
        • Audio Sciences: Recording Arts and Sciences, Master of Arts
      • Doctor of Musical Arts (DMA)
        • Composition, Doctor of Musical Arts
        • Performance, Doctor of Musical Arts -​ Choral Conducting specialization
        • Performance, Doctor of Musical Arts -​ Guitar specialization
        • Performance, Doctor of Musical Arts -​ Historical Performance Instruments specialization
        • Performance, Doctor of Musical Arts -​ Orchestral Conducting specialization
        • Performance, Doctor of Musical Arts -​ Orchestral Instruments specialization
        • Performance, Doctor of Musical Arts -​ Organ specialization
        • Performance, Doctor of Musical Arts -​ Piano specialization
        • Performance, Doctor of Musical Arts -​ Voice specialization
        • Performance, Doctor of Musical Arts -​ Wind Conducting specialization
      • Performer’s Certificate (PC)
        • Guitar, Performer's Certificate
        • Orchestral Instruments, Performer's Certificate
        • Organ, Performer's Certificate
        • Piano, Performer's Certificate
        • Voice, Performer's Certificate
      • Graduate Performance Diploma (GPD)
      • Artist’s Diploma (AD)
    • Extension Study
      • Music Education Certification -​ Instrumental
      • Music Education Certification -​ Vocal
  • Nitze School of Advanced International Studies
    • Degrees and Certificates
      • International Studies, Doctor of Philosophy
      • International Affairs, Doctor of
      • European Public Policy, Master of Arts
      • Global Policy, Master of Arts
      • Global Risk, Master of Arts (On-​site)
      • Global Risk, Master of Arts (Online)
      • International Affairs, Master of Arts
      • International Economics and Finance, Master of Arts
      • International Relations, Master of Arts
      • International Studies, Master of Arts
      • International Public Policy, Master of
      • Strategy, Cybersecurity, and Intelligence, Master of Arts
      • Sustainable Energy, Master of Arts (Online)
      • Chinese and American Studies, Hopkins-​Nanjing Center Certificate
      • Dual Degrees and Exchange Programs
      • Graduate Certificates
      • International Studies, Diploma
    • Policies and Resources
      • Academic Integrity
      • Academic Policies and Resources
      • Student Life
    • School Leadership and Key Contacts
  • School of Education
    • Academic and Student Policies
      • Academic and Student Conduct Policies
      • Academic Standards
      • Grading System and Academic Records
      • Grievances and Complaints
    • Admission
    • Graduation
    • Programs
      • Doctoral Programs
        • Education (Online), EdD
        • Education, PhD
      • Master's Programs
        • Counseling, Master of Science
        • Education, Master of Science
          • Education, Master of Science – Digital Age Learning and Educational Technology (Online)
          • Education, Master of Science -​ Educational Studies
          • Education, Master of Science -​ Gifted Education
        • Education Policy, Master of Science
        • Health Professions (Online), Master of Education
        • Learning Design and Technology, Master of Education
        • Special Education, Master of Science
        • Teaching Professionals, Master of Education
      • Post Master's Certificates
        • Applied Behavior Analysis, Post–Master’s Certificate
        • Evidence-​Based Teaching in the Health Professions, Post–Master’s Certificate
    • Centers &​ Institutes
    • Scholarships
    • State Authorization of Distance Education (NC-​SARA)
  • School of Medicine
    • General Information
      • Conduct in Teacher/​Learner Relationships (Learner Treatment Policy)
      • Lectureships and Visiting Professorships
      • Loan Funds
      • Medical Student Advising
      • Named Professorships
      • Office of Medical Student Affairs
      • Scholarships
      • Student Research Scholarships and Awards
      • Tuition
      • Tuition and Other Fees
      • Young Investigators’ Day
    • Policies
    • Graduate Programs
      • Anatomy Education, MS
      • Applied Health Sciences Informatics, MS
      • Biochemistry, Cellular and Molecular Biology, PhD
      • Biological Chemistry, PhD
      • Biomedical Engineering, PhD
      • Cellular and Molecular Medicine, MS
      • Cellular and Molecular Medicine, PhD
      • Cellular and Molecular Physiology, PhD
      • Clinical Anaplastology, MS
      • Clinical Informatics, Post-​Baccalaureate Certificate
      • Cross-​Disciplinary Program in Graduate Biomedical Sciences, PhD
      • Functional Anatomy and Evolution, PhD
      • Health Sciences Informatics, MS
      • Health Sciences Informatics, PhD
      • History of Medicine, MA (On-​site)
      • History of Medicine, MA (Online)
      • History of Medicine, PhD
      • History of Medicine, Post-​Baccalaureate Certificate (Online)
      • Human Genetics and Genomics, PhD
      • Immunology, PhD
      • Medical and Biological Illustration, MA
      • Medical Physics, MS
      • Medical Physics, PhD
      • Medical Physics, Post-​Baccalaureate Certificate
      • Molecular Biophysics, PhD
      • Neuroscience, PhD
      • Pathobiology, PhD
      • Pharmacology and Molecular Sciences, PhD
    • Medical Program
      • Doctor of Medicine, MD
      • MD-​MBA, Combined Degree
      • MD-​PhD, Combined Degree
      • Subject Areas
        • Anesthesiology and Critical Care Medicine
        • Biological Chemistry
        • Biomedical Engineering
        • Biophysics and Biophysical Chemistry
        • Cell Biology
        • Department of Genetic Medicine
        • Dermatology
        • Emergency Medicine
        • Epidemiology
        • Functional Anatomy and Evolution
        • Gynecology and Obstetrics
        • Health Sciences Informatics
        • History of Medicine
        • Medicine
        • Molecular and Comparative Pathobiology
        • Molecular Biology and Genetics
        • Multi-​Department Courses
        • Neurology
        • Neuroscience
        • Oncology
        • Ophthalmology
        • Pathology
        • Pediatrics
        • Physical Medicine and Rehabilitation
        • Physiology, Pharmacology and Therapeutics
        • Psychiatry and Behavioral Sciences
        • Public Health
        • Radiation Oncology and Molecular Radiation Sciences
        • Radiology and Radiological Science
        • Surgery
    • Postdoctoral Fellows
  • School of Nursing
    • Admission
    • Advising
    • Certificates
      • Healthcare Organizational Leadership, Post-​Master’s Certificate
      • Nursing Education, Post-​Master's Certificate
      • Psychiatric Mental Health Nurse Practitioner, Post-​Master's Certificate
    • Doctoral Degrees
      • Doctor of Nursing Practice, Advanced Practice Track
        • Adult-​Gerontological Acute Care Nurse Practitioner, DNP Advanced Practice Track
        • Adult-​Gerontological Critical Care Clinical Nurse Specialist, DNP Advanced Practice Track
        • Adult-​Gerontological Health Clinical Nurse Specialist, DNP Advanced Practice Track
        • Adult-​Gerontological Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Family Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Nurse Anesthesia, DNP Advanced Practice Track
        • Pediatric Critical Care Clinical Nurse Specialist, DNP Advanced Practice Track
        • Pediatric Dual Primary/​Acute Care Nurse Practitioner, DNP Advanced Practice Track
        • Pediatric Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Psychiatric Mental Health Nurse Practitioner, DNP Advanced Practice Track
      • Doctor of Nursing Practice: Post Master's Track
      • Nursing, Doctor of Philosophy
      • Doctor of Nursing Practice (DNP): Advanced Practice Track/​Doctor of Philosophy in Nursing (PhD) Dual Degree
    • Dual Degrees
      • DNP Post Master's/​MBA Dual Degree
      • DNP Post Master's/​MPH Dual Degree
      • Healthcare Organizational Leadership, MSN/​MBA, Dual Degree
    • Financial Aid
    • Master's Degrees
      • Entry into Nursing, Master of Science in Nursing
      • Healthcare Organizational Leadership Track, Master of Science in Nursing
    • Online Prerequisites for Health Professions
    • Policies
      • Academic Integrity Policy
      • Academic Standards for Progression
      • Administrative Leave
      • Absence and Attendance Policy
      • Canvas and SON IT Help
      • Clinical Placements
      • Clinical Warnings
      • Complaint/​Grievance Policy
      • Compliance
      • Course Policies
      • Criminal Conduct/​Background Check Policies
      • Drug Testing Policy
      • Email Policy
      • Examination Policy
      • Grading Policy
      • Health Insurance for Students
      • Incomplete Coursework
      • Independent Study Policy
      • Leave of Absence
      • Letters of Recommendation
      • NCLEX
      • Non-​Degree-​Seeking Students
      • Notification of Missed Clinical Time
      • Pet Guidelines
      • Printing and Copying
      • Professional Attire Policy
      • Professional Ethics Policy
      • Registration Policies and Procedures
      • Religious Accommodation
      • Social Media Guidelines
      • Student Code of Conduct
      • Technical Standards for Admission and Graduation
      • Transcripts and Enrollment Verifications
      • Transfer of Graduate Credit
      • Withdrawal Policy
    • Student Accounts
    • Tuition and Fees
  • Whiting School of Engineering
    • Full-​time, On-​campus Undergraduate and Graduate Programs (Homewood)
      • Zanvyl Krieger School of Arts and Sciences &​ Whiting School of Engineering Full-​Time, On-​Campus Undergraduate Policies
      • Whiting School of Engineering Graduate Policies
        • Academic Policies
        • Admissions and Finances
        • Graduate-​Specific Policies
        • Student Life
          • International Graduate Students
      • Departments, Program Requirements, and Courses
        • Applied Mathematics and Statistics
          • Applied Mathematics and Statistics, Bachelor of Arts
          • Applied Mathematics and Statistics, Bachelor of Science
          • Applied Mathematics and Statistics, Master of Science in Engineering
          • Applied Mathematics and Statistics, Minor
          • Applied Mathematics and Statistics, PhD
          • Data Science, Master of Science in Engineering
          • Financial Mathematics, Master of Science in Engineering
        • Biomedical Engineering
          • Bioengineering Innovation and Design, Master of Science in Engineering
          • Biomedical Engineering, Bachelor of Science
          • Biomedical Engineering, Master of Science in Engineering
          • Biomedical Engineering, PhD through the School of Medicine
        • Center for Leadership Education
          • Accounting and Financial Management, Minor
          • Engineering Management, Master of Science
          • Global Innovation and Leadership Through Engineering, Master of Science
          • Leadership Studies, Minor
          • Marketing and Communications, Minor
          • Professional Communication Program
          • Professional Development Program
          • W.P. Carey Entrepreneurship and Management, Minor
        • Chemical and Biomolecular Engineering
          • Chemical and Biomolecular Engineering, Bachelor of Science
          • Chemical and Biomolecular Engineering, Master of Science in Engineering
          • Chemical and Biomolecular Engineering, PhD
        • Civil &​ Systems Engineering
          • Civil Engineering, Bachelor of Science
          • Civil Engineering, Master of Science in Engineering (MSE)
          • Civil Engineering, Minor
          • Civil and Systems Engineering, PhD
          • Systems Engineering, Bachelor of Science
          • Systems Engineering, Master of Science
          • Systems Engineering, Minor
        • Computational Medicine
          • Computational Medicine, Minor
        • Computer Science
          • Computer Science, Bachelor of Arts
          • Computer Science, Bachelor of Science
          • Computer Science, Master of Science in Engineering
          • Computer Science, Minor
          • Computer Science, PhD
        • Doctor of Engineering
          • Engineering, Doctor of Engineering
        • Electrical and Computer Engineering
          • Computer Engineering, Bachelor of Science
          • Electrical and Computer Engineering, Master of Science in Engineering
          • Electrical and Computer Engineering, PhD
          • Electrical Engineering, Bachelor of Science
          • Energy, Minor
        • Environmental Health and Engineering
          • Engineering for Sustainable Development, Minor
          • Environmental Engineering, Bachelor of Science
          • Environmental Engineering, Minor
          • Environmental Engineering, PhD
          • Environmental Health and Engineering, Master of Arts
          • Environmental Health and Engineering, Master of Science
          • Environmental Health and Engineering, Master of Science in Engineering
          • Environmental Sciences, Minor
          • Occupational and Environmental Hygiene, Master of Science
        • General Engineering
          • General Engineering, Bachelor of Arts
        • Information Security Institute
          • Security Informatics, Master of Science
          • Security Informatics, Master of Science/​Applied Mathematics and Statistics, Master of Science in Engineering Dual Master's Program
          • Security Informatics, Master of Science/​Computer Science, Master of Science in Engineering Dual Master's Program
        • Materials Science and Engineering
          • Materials Science and Engineering, Bachelor of Science
          • Materials Science and Engineering, Master of Science in Engineering
          • Materials Science and Engineering, PhD
        • Mechanical Engineering
          • Engineering Mechanics, Bachelor of Science
          • Mechanical Engineering, Bachelor of Science
          • Mechanical Engineering, Master of Science in Engineering
          • Mechanical Engineering, PhD
        • NanoBioTechnology
        • Robotics and Computational Sensing
          • Computer Integrated Surgery, Minor
          • Robotics, Master of Science in Engineering
          • Robotics, Minor
      • Multi-​School Programs of Study
        • Business, Minor
        • Peabody-​Homewood Double Degree Program
        • Space Science and Engineering
    • Part-​Time, Online Graduate Programs (Engineering for Professionals)
      • Academic Policies
        • Academic Calendar
        • Academic Regulations
        • Registration Policies
        • Tuition and Fees
      • Admission Requirements
      • Applied and Computational Mathematics
        • Applied and Computational Mathematics, Graduate Certificate
        • Applied and Computational Mathematics, Master of Science
        • Applied and Computational Mathematics, Post-​Master’s Certificate
      • Applied Biomedical Engineering
        • Applied Biomedical Engineering, Graduate Certificate
        • Applied Biomedical Engineering, Master of Science
        • Applied Biomedical Engineering, Post-​Master’s Certificate
      • Applied Physics
        • Applied Physics, Master of Science
        • Applied Physics, Post-​Master’s Certificate
      • Artificial Intelligence
        • Artificial Intelligence, Graduate Certificate
        • Artificial Intelligence, Master of Science
      • Chemical and Biomolecular Engineering
        • Chemical and Biomolecular Engineering, Master of Chemical and Biomolecular Engineering
      • Civil Engineering
        • Civil Engineering, Graduate Certificate
        • Civil Engineering, Master of Civil Engineering
      • Computer Science
        • Computer Science, Graduate Certificate
        • Computer Science, Master of Science
        • Computer Science, Post-​Master’s Certificate
      • Cybersecurity
        • Cybersecurity, Graduate Certificate
        • Cybersecurity, Master of Science
        • Cybersecurity, Post-​Master’s Certificate
      • Data Analytics and Engineering
        • Data Analytics and Engineering, Master of Science
      • Data Science
        • Data Science, Graduate Certificate
        • Data Science, Master of Science
        • Data Science, Post-​Master’s Certificate
      • Electrical and Computer Engineering
        • Electrical and Computer Engineering, Graduate Certificate
        • Electrical and Computer Engineering, Master of Science
        • Electrical and Computer Engineering, Post-​Master’s Certificate
      • Engineering Management
        • Engineering Management, Graduate Certificate
        • Engineering Management, Master of Engineering Management
      • Environmental Engineering, Science, Management, and Sustainability Programs
        • Climate, Energy, and Environmental Sustainability, Graduate Certificate
        • Climate, Energy, and Environmental Sustainability, Master of Science
        • Environmental Engineering
          • Environmental Engineering, Graduate Certificate
          • Environmental Engineering, Master of Environmental Engineering
          • Environmental Engineering, Post-​Master’s Certificate
        • Environmental Engineering and Science
          • Environmental Engineering and Science, Graduate Certificate
          • Environmental Engineering and Science, Master of Science
          • Environmental Engineering and Science, Post-​Master’s Certificate
        • Environmental Planning and Management
          • Environmental Planning and Management, Graduate Certificate
          • Environmental Planning and Management, Master of Science
          • Environmental Planning and Management, Post-​Master’s Certificate
      • Financial Mathematics
        • Financial Mathematics, Master of Science
        • Financial Risk Management, Graduate Certificate
        • Quantitative Portfolio Management, Graduate Certificate
        • Securitization, Graduate Certificate
      • Healthcare Systems Engineering
        • Healthcare Systems Engineering, Master of Science
      • Industrial and Operations Engineering
        • Industrial and Operations Engineering, Master of Science
      • Information Systems Engineering
        • Information Systems Engineering, Graduate Certificate
        • Information Systems Engineering, Master of Science
        • Information Systems Engineering, Post-​Master’s Certificate
      • Materials Science and Engineering
        • Materials Science and Engineering, Master of Science
      • Mechanical Engineering
        • Mechanical Engineering, Master of Science
        • Mechanical Engineering, Post-​Master’s Certificate
      • Occupational and Environmental Hygiene
        • Occupational and Environmental Hygiene, Master of Science
      • Robotics and Autonomous Systems
        • Robotics and Autonomous Systems, Master of Science
      • Space Engineering
        • Space Engineering, Master of Science
        • Space Engineering, Post-​Master's Certificate
      • Systems Engineering
        • Systems Engineering, Graduate Certificate
        • Systems Engineering, Master of Science
        • Systems Engineering, Master of Science in Engineering (ABET-​accredited)
        • Systems Engineering, Post-​Master’s Certificate
  • Zanvyl Krieger School of Arts and Sciences
    • Full-​time, On-​campus Undergraduate and Graduate Programs (Homewood)
      • Zanvyl Krieger School of Arts and Sciences &​ Whiting School of Engineering Full-​Time, On-​Campus Undergraduate Policies
      • Krieger School of Arts &​ Sciences Graduate Policies
        • Academic Policies
        • Admissions and Finances
        • Graduate-​Specific Policies
        • Student Life
          • International Graduate Students
      • Departments, Program Requirements, and Courses
        • Anthropology
          • Anthropology, Bachelor of Arts
          • Anthropology, Minor
          • Anthropology, PhD
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  • Neuroscience, PhD
  • Overview
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Overview

The Department of Neuroscience offers an interdisciplinary program designed to train doctoral students for independent research and teaching in neuroscience. It is the goal of the program to ensure that candidates for the PhD and MD/PhD degrees obtain a background covering molecular, cellular, systems, and cognitive approaches to neuroscience, as well as receive training that brings them to the forefront of research in their particular area of interest. The program also ensures that students obtain broad training in the neurosciences. Our curriculum spans the breadth of modern neuroscience, from molecular/cellular underpinnings to systems/cognitive integration and offers a rich training experience. Students enter the program from different backgrounds and join laboratories working on different disciplines; therefore, the program is tailored to fit the needs of individual students. Courses are designed so that students have ample time to become involved in laboratory research in the first year. Most students begin their thesis research at the beginning of their second year.

For more information, please visit The Solomon H. Snyder Department of Neuroscience webpage.

Training Facilities

The Neuroscience Training Program is based in the Solomon S. Snyder Department of Neuroscience. The Training Program includes approximately 120 faculty and laboratory facilities located in the Department of Neuroscience as well as several other basic and clinical science departments closely associated with the Neuroscience Department. Modern state-of-the-art facilities for research in molecular biology, neurophysiology, pharmacology, biochemistry, cell biology, and morphology are available. The Mind/Brain Institute, located on the Homewood Campus of the University, is a group of laboratories devoted to the investigation of the neural mechanisms of higher mental function and particularly to the mechanisms of perception. All of the faculty in the Mind/Brain Institute are members of the Neuroscience Graduate Program.

Neuroscience trainees may also elect to conduct research in laboratories at the Howard Hughes Medical Institute’s Janelia Research Campus (19700 Helix Drive, Ashburn, Virginia 20147). The Janelia Research Campus provides research opportunities in small labs, encourages collaboration across different laboratories and research topics, and offers access to developing tools and technologies. Johns Hopkins University is certified to operate elements of the program at this site by the State Council of Higher Education for Virginia.

Financial Aid

Candidates accepted into the Program are offered full support providing payment of tuition, health, dental and vision benefits as well as a stipend for the duration of their studies. For more details regarding financial aid opportunities, please visit Financial Aid | Johns Hopkins Medicine. All entering and first-year students are encouraged to apply for individual fellowships such as those sponsored by the National Science Foundation and the Howard Hughes Medical Institute.

Vivien Thomas PhD Scholars at JHU

The Vivien Thomas Scholars Initiative (VTSI) is an endowed fellowship program at Johns Hopkins for PhD students in STEM fields. Vivien Thomas Scholars are selected from a range of eligible institutions, including Historically Black Colleges and Universities (HBCUs), community colleges across the country, and regional institutions in Maryland, Virginia, and the District of Columbia. In addition to the interview and selection process for the VTSI program, they must also be accepted into a Johns Hopkins PhD STEM program through the regular, rigorous and competitive application process. Scholars who have been accepted into both a PhD STEM program and the VTSI receive the financial support needed to ensure their success, including up to six years of full tuition support, a generous stipend, health insurance, and other benefits, along with significant mentorship, research, professional development, and community-building opportunities. More information about the VTSI program is available at this link. 

Admission Requirements

Note: PhD applications are submitted via the School of Medicine's application system, and the deadline is usually in the first week of December. The Neuroscience program does not require GRE scores. The Neuroscience website has up-to-date information on the ‘how to apply’ page.

Applicants are expected to have received a B.S. or B.A. prior to enrolling in the graduate program. Laboratory research experience prior to enrollment is also desirable. We are most enthusiastic about applicants who have taken full advantage of the opportunities available at their undergraduate institution and through other summer or postbac experiences. If you have research experience, please describe your research in your Statement of Interest and Career Objectives and indicate the number of months engaged in full-time and part-time research on your CV. Students who do well in our program typically have a strong academic foundation in areas of biological or physical sciences. Some of the courses that prepare students well include general biology, neuroscience, mathematics through calculus, general physics, general chemistry, organic chemistry, statistics, engineering, or computer science.

Combined MD/PhD Program 

A subset of the current predoctoral trainees in the Neuroscience Program are candidates for both PhD and MD degrees. Applications for admission to the combined program are considered by the MD/PhD Committee of the School of Medicine. Application forms for the School of Medicine contain a section requesting information relevant to graduate study. Applicants interested in the combined MD/PhD program should complete this section and indicate specifically their interest in the Neuroscience Training Program.  Information about applying can be found at the Hopkins MSTP admissions page.

Program Requirements

Courses

During the first two years in the Neuroscience Training Program (NTP), students are required to take several graduate level core courses as well as advanced electives that provide rigorous training in principles of neuroscience research. In addition, students in the first year attend Quantitative Analysis boot camp and Neuroscience Research Symposia in the first few weeks after joining the program.  

Course List
Code Title Credits
Required Courses
ME.440.811Neuroscience Cognition I4.5
ME.440.812Neuroscience Cognition II4.5
ME.440.819Rigor, Reproducibility, and Responsibility in Science 12
ME.440.801Readings in Neuroscience (Journal Club) 21
or ME.440.810 Readings In Systems Neuroscience
ME.440.802Current Topics in Neuroscience (Research Seminar) 31
or AS.080.631 Bodian Seminar Series
ME.440.820Circuits and Brain Disorders2
AS.200.659Quantitative Methods for Brain Sciences3
or ME.440.825 Quantitative Neurogenomics
PH.140.615Statistics for Laboratory Scientists I4
PH.140.616Statistics for Laboratory Scientists II4
ME.440.803Teaching in Neuroscience1
ME.440.724Neuroscience Career Skills1
ME.440.730Submitting Your First Paper0.5
ME.440.823Grant Writing Skills1
ME.440.800Neuroscience Research 41 - 18
Neuroscience Elective Courses
A minimum of 3 pre-approved electives (with at least one primarily focused on a neuroscience topic) are required. Neuroscience electives are listed below. A full listing of pre-approved courses can be found in the NTP program guidelines.
AS.080.620Theoretical and Computational Neuroscience3
ME.440.822Computational Principles of Biological Vision3
AS.080.310Synaptic Function and Plasticity3
ME.440.709Neuropharmacology1.5
ME.440.715Trends in the Neurobiology of Aging0.5
ME.440.818Bioenergetics, Neuroplasticity and Brain Health1
ME.440.711Cellular and Molecular Basis of Neural Development I: Neuronal Differentiation1.5
ME.440.705Cellular and Molecular Basis of Neural Development II1.5
ME.440.723Writing About the Brain3
ME.440.707Molecular Mechanisms in Synaptic Transmission2
EN.580.625Structure and Function of the Auditory and Vestibular Systems3
ME.440.824Cell Physiology of Visual and Olfactory Transductions1
1

Completion of this course fulfills the SOM Ethics Requirement, and needs to be competed in fall year 1 and repeated fall year 5.

2

Required fall and spring semesters in years 1, 2 and 3

3

Required fall and spring semesters in years 1 and 2 

4

Required every semester until graduation for full-time enrollment

Laboratory Rotations

Lab rotations are required for students who join the program on a Rotation track. These laboratory rotations expose the student to a variety of current research techniques in neuroscience and provide an opportunity for the student to select a laboratory in which to conduct dissertation research. Scheduling of the three rotations is adjusted to make the most convenient schedule for each student, and require a rotation report submitted to the program at the end of each rotation. The rotations are usually completed by the end of the first year. Each rotation track student will complete 3 research rotations prior to selecting a thesis laboratory. Declaring a thesis laboratory before the completion of 3 rotations is an option and requires steering committee approval. A fourth rotation may be completed if necessary/desired. Rotations at the Janelia Research Campus typically happen during the summer. Generally, each research rotation will be about 3 months in length, with the exception of summer rotations. A full-time summer rotation (July-August) will be about 2 months in length. It is critical for rotating students to consult with rotation mentors (PIs) about available funding for the entire training period before a rotation is engaged.

Students in the Direct Admit track do not complete rotations but rather join their sponsor’s lab upon matriculation.

Annual Neuroscience Retreat

Attendance at the Annual Neuroscience Retreat is required for all students.

Monthly Lab Talks

Once a month, the faculty, postdoctoral fellows, and students from one laboratory present and discuss the ongoing research in that laboratory. This provides an informal setting to discuss research being conducted in the laboratories of the Neuroscience Training Program and gives advanced graduate students and postdoctoral fellows a forum for presenting their work. Attendance at the monthly Neuroscience Lab Talks is required for all students.

Annual IDP

Students must complete annual meetings with their faculty advisor on the student's academic progress and professional development plan, also known as the student's Individual Development Plan (IDP).

Residency requirements 

A minimum residency of two consecutive semesters is required. During the course of graduate study, the student must successfully complete all program requirements.

Doctor of Philosophy Board Oral  (DBO) Examination

A University-mandated Doctor of Philosophy Board Oral  (DBO) Examination must be completed by the second year of study.  The DBO Examination tests the breadth and depth of the graduate student’s scientific knowledge and readiness to begin thesis research.

Thesis Advisory Committee Meetings

Following the DBO, the first Thesis Committee meeting is called the Thesis Proposal Defense. The student must prepare a thesis proposal that follows the guidelines and format of the NIH Pre-doctoral NRSA Application. The proposal should be well-formulated and presented in sufficient detail that it can be evaluated for both its research training potential and scientific merit. It is important that it be developed in collaboration with the thesis advisor, but it must be written entirely by the student.

Annual thesis committee meetings are held in year 3 and 4, and every 6 months thereafter, until such time as the thesis committee evaluates that the student is ready to write their doctoral dissertation and receive permission to write. Submission of a manuscript is a requirement for the committee to grant permission to write.

Dissertation Defense

The dissertation is based on the student’s novel research and must be approved by all four members of the Thesis Advisory Committee before the student can graduate. This happens during the Dissertation Defense meeting. Following a successful dissertation defense, the thesis is submitted to the Sheridan Library. The thesis advisor and second reader submit a reader's letter approving the dissertation, via the Neuroscience program. The complete graduation package needs to be approved by the registrar. A public seminar of this thesis work is a graduation requirement. 

Core competencies in Neuroscience

1.  Basic knowledge of neuroscience principles

All graduates should be able to:

  • Explain how neurons use passive and active membrane properties to receive, process, and transmit signals to post-synaptic cells. 
  • Explain how neurons are connected in circuits and refined by experience.
  • Demonstrate a fundamental understanding of the cellular and molecular specializations that support nervous system function.
  • Demonstrate sufficient understanding of neuroanatomy to describe neural circuits and how they contribute to nervous system function.
  • Explain how the nervous system develops.
  • Identify and discuss common disorders of the nervous system, their biological basis, & current therapies.
  • Grasp the fundamental approaches used by theoretical neuroscience to understand how the brain solves computational problems.
  • Demonstrate familiarity with common methodologies used in experimental neuroscience.
  • Explain biochemical and cell biological processes important for brain function.

2.  Critical thinking

All graduates should be able to:

  • Analyze primary literature and identify strengths and shortcomings of the methods employed.
  • Construct testable hypotheses and design experiments to challenge these hypotheses.

3.  Quantitative analysis

All graduates should be able to: 

  • Appropriately process and apply analytical techniques and statistical tests to their data.
  • Create figures that effectively communicate results.
  • Critically evaluate and interpret quantitative data.
  • Identify and use computational tools in their research.

4.  Effective written/oral communication skills

All graduates should be able to:

  • Organize their oral and written scientific communications to effectively transmit: 1. significance of topic, 2. relevant background material to place the topic in context, and 3. knowledge gap to be addressed.
  • Orally present scientific material in a clear and effective manner.

5.  Career-specific skills

All graduates should be able to:

  • Develop leadership and management skills and cultural awareness required to translate their neuroscience Ph.D. into a successful career.
  • Leverage discussions with their advisor, advisory committee,  IDP, Career Development Office programs, and other available resources for advocacy, education, outreach and career path considerations.

6.  Self-motivated learning/scientific inquiry

All graduates should:

  • Independently explore and assimilate existing literature in their field of interest.
  • Be able to identify and engage expert guidance when needed.

7.  Discipline-specific research skills

All graduates should be able to:

  • Develop hypothesis driven research questions founded on own their current studies.
  • Conduct discipline-specific experimental techniques with appropriate controls and analysis.
  • Troubleshoot and solve emergent problems.
  • Explain how their studies may impact human health.

8.  Citizenship

All graduates should demonstrate:

  • A fundamental understanding of research ethics.
  • The ability to work collaboratively with others.
  • Leadership skills necessary to effectively train and supervise others.
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