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Neuroscience, PhD

School of Medicine

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  • Explore our Programs
  • University-​wide Policies and Information
    • Academic Policies and Information
      • Academic Calendar
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      • Animal Care and Use Program
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      • Student Leave of Absence Policy
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    • Admission and Aid
      • Tuition, Fees, and Cost of Attendance
        • Financial Aid
    • Higher Education Act Disclosures
      • General Institutional Information
      • Health and Safety Information
      • Student Financial Assistance Information
    • Office of Institutional Equity
      • Discrimination and Harassment Policy and Procedures
      • Equal Opportunity and Title IX Notice
      • Sexual Misconduct Policy and Procedures
    • Rights, Privileges, and Responsibilities
      • Academic Grievance Policy: Students and Postdoctoral Fellows
      • New Child Accommodations for Full-​Time Graduate Students and Postdoctoral Trainees
      • Personal Relationships Policy
      • Photography and Film Rights Policy
      • Student Conduct Code
      • Student Disability Services (SDS)
      • Student Health
    • Veterans Affairs
  • Bloomberg School of Public Health
    • Academic Calendar
    • Admission
    • CEPH Requirements
    • Departments
      • Department of Biochemistry and Molecular Biology
        • Biochemistry and Molecular Biology, MHS
        • Biochemistry and Molecular Biology, ScM
        • Biochemistry and Molecular Biology, PhD
        • Non-​Degree Training
      • Department of Biostatistics
        • Biostatistics, MHS
        • Biostatistics, ScM
        • Biostatistics, PhD
      • Department of Environmental Health and Engineering
        • Environmental Health, MHS
        • Environmental Health, SCM
        • Toxicology for Human Risk Assessment, MS
        • Environmental Health, PhD
        • Non-​Degree Training
      • Department of Epidemiology
        • Epidemiology, MHS
        • Epidemiology, ScM
        • Epidemiology, PhD
        • Non-​Degree Training
      • Department of Health, Behavior and Society
        • Health Education and Health Communication, MSPH
        • Genetic Counseling, ScM
        • Health, Behavior, and Society, MHS
        • Health, Behavior and Society, PhD
        • Non-​Degree Training
      • Department of Health Policy and Management
        • Health Administration, MHA
        • Health Economics and Outcomes Research, MHS
        • Health Finance and Management, MHS
        • Health Policy, MSPH
        • Health Policy and Management, PhD
        • Health Policy and Management, DrPH (Tsinghua)
        • Non-​Degree Training
      • Department of International Health
        • Global Health Economics, MHS
        • International Health, MSPH
        • International Health, MSPH, Human Nutrition-​Dietitian
        • International Health, MA/​MSPH
        • International Health, PhD
        • Non-​Degree Training
      • Department of Mental Health
        • Mental Health, MHS
        • Mental Health, PhD
        • Non-​Degree Training
      • Department of Molecular Microbiology &​ Immunology
        • Molecular Microbiology &​ Immunology, MHS
        • Molecular Microbiology &​ Immunology, ScM
        • Molecular Microbiology &​ Immunology, PhD
        • Non-​Degree Training
      • Department of Population, Family and Reproductive Health
        • Population, Family and Reproductive Health, MHS
        • Population, Family and Reproductive Health, MHS Online
        • Population, Family and Reproductive Health, MSPH
        • Population, Family and Reproductive Health, PhD
      • Doctor of Public Health (DrPH)
      • Graduate Training Programs in Clinical Investigation
        • Graduate Training Programs in Clinical Investigation, MHS
        • Graduate Training Programs in Clinical Investigation, PhD
        • Graduate Training Programs in Clinical Investigation, ScM
      • Master of Arts in Public Health Biology
      • Master of Bioethics
      • Master of Public Health Program
        • DNP/​MPH
        • DVM/​MPH
        • JD/​MPH
        • LLM/​MPH
        • MBA/​MPH with China Europe International Business School
        • MD/​MPH
        • MPH/​MBA
        • MSW/​MPH
      • MAS-​Office
        • Master of Applied Science in Community-​Based Primary Health Care Programs in Global Health
        • Master of Applied Science in Humanitarian Health
        • Master of Applied Science in Patient Safety and Healthcare Quality
        • Master of Applied Science in Population Health Management
        • Master of Applied Science in Spatial Analysis for Public Health
      • Residency Programs
        • General Preventive Medicine Residency Program
        • Occupational and Environmental Medicine Residency
    • Certificates
      • Adolescent Health, Certificate
      • Bioethics, Certificate
      • Climate and Health, Certificate
      • Clinical Trials, Certificate
      • Community-​Based Public Health, Certificate
      • Demographic Methods, Certificate
      • Environmental and Occupational Health, Certificate
      • Epidemiology for Public Health Professionals, Certificate
      • Evaluation: International Health Programs, Certificate
      • Food Systems, the Environment &​ Public Health, Certificate
      • Gender and Health, Certificate
      • Gerontology, Certificate
      • Global Digital Health, Certificate
      • Global Health, Certificate
      • Global Health Practice, Certificate
      • Health and Human Rights, Certificate
      • Health Communication, Certificate
      • Health Disparities and Health Inequality, Certificate
      • Health Education, Certificate
      • Health Finance and Management, Certificate
      • Healthcare Epidemiology and Infection Prevention and Control, Certificate
      • Humane Sciences and Toxicology Policy, Certificate
      • Humanitarian Health, Certificate
      • Implementation Science and Research Practice, Certificate
      • Injury and Violence Prevention, Certificate
      • International Healthcare Management and Leadership, Certificate
      • Leadership for Public Health and Healthcare, Certificate
      • Lesbian, Gay, Bisexual, Transgender, and Queer (LGBTQ) Public Health, Certificate
      • Maternal and Child Health, Certificate
      • Mental Health Policy, Economics and Services, Certificate
      • Pharmacoepidemiology and Drug Safety, Certificate
      • Population and Health, Certificate
      • Population Health Management, Certificate
      • Product Stewardship for Sustainability, Certificate
      • Public Health Advocacy, Certificate
      • Public Health Economics, Certificate
      • Public Health Informatics, Certificate
      • Public Health Practice, Certificate
      • Public Health Preparedness, Certificate
      • Public Health Training Certificate for American Indian Health Professionals
      • Public Mental Health Research, Certificate
      • Quality, Patient Safety, and Outcomes Research, Certificate
      • Quantitative Methods in Public Health, Certificate
      • Rigor, Reproducibility and Responsibility in Scientific Practice, Certificate
      • Risk Sciences and Public Policy, Certificate
      • Spatial Analysis for Public Health, Certificate
      • Training Certificate in Public Health
      • Tropical Medicine, Certificate
      • Vaccine Science and Policy, Certificate
    • Policies
      • Academic
        • Academic Ethics Code
        • Compliance Line
        • Grade Appeal Policy
        • Grading System
        • Graduation Policy
        • Interdivisional Registration
        • Multi-​Term Course Policy
        • Post-​Doctoral Fellow Student Status
        • Student Grievance Policy
        • Voluntary Leave of Absence Policy
      • Research
        • Animal Research
        • Human Subjects Research
        • Worker's Compensation
  • Carey Business School
    • Admission
      • Master’s Programs
      • Certificate Programs
      • Graduate Degree Requirements
      • International Student Admission Policy
      • Verification of Credentials
      • Other Admission Policies
    • Degrees and Certificates
      • Business Administration (Flexible), MBA
      • Business Administration (Full Time), MBA
      • Business Analytics and Risk Management (Part Time), Master of Science
      • Business Analytics and Risk Management, Master of Science
      • Design Leadership, MBA/​MA Dual Degree
      • Finance (Part Time), Master of Science
      • Finance, Master of Science
      • Financial Management, Graduate Certificate
      • Financial Management, Graduate Certificate, Investments, Graduate Certificate, Applied Economics, MS
      • Health Care Management (Part Time), Master of Science
      • Health Care Management, Master of Science
      • Information Systems, Master of Science
      • Investments, Graduate Certificate
      • Leadership Development Program, Graduate Certificate
      • Marketing (Part Time), Master of Science
      • Marketing, Master of Science
      • MBA/​Applied Economics, MS Dual Degree
      • MBA/​Biotechnology, MS Dual Degree
      • MBA/​Communication, MA Dual Degree
      • MBA/​DNP Dual Degree
      • MBA/​Government, MA Dual Degree
      • MBA/​Healthcare Organizational Leadership, MSN Dual Degree
      • MBA/​JD Dual Degree
      • MBA/​MA in International Relations
      • MBA/​MD Dual Degree
      • MBA/​MPH Dual Degree
      • Real Estate and Infrastructure (Part Time), Master of Science
      • Real Estate and Infrastructure, Master of Science
      • Business, Minor
    • Policies and Resources
      • Academic Calendar
      • Academic Ethics Policy
      • Academic Progress and Standards
      • Changing Degree Program
      • Grading Policy
      • Graduation
      • Attendance Policy
      • Leave of Absence
      • Registration
      • Student Accounts
      • Transfer of Graduate Credit
      • Waiver Exams
  • Peabody Institute
    • General Information, Procedures and Regulations
      • Introduction and Nomenclature
      • Mission
      • Accreditation
      • Links
      • Honor Societies
    • Procedural Information
      • Applicability
      • Studio Assignments
      • Course Numbering
      • Large Ensemble Participation
      • Competitions
      • Recitals
      • Academic Advising
      • Inter-​Institutional Academic Arrangements
      • Study Abroad Program
      • Outside Instruction and Public Performance
    • Academic Regulations
      • Applicability
      • Academic Code of Conduct
      • Program Classification, Status, and Credit Limits
      • Sources of Credit
      • Grading System and Regulations
      • Dean's List Criteria
      • Academic Standing
      • Registration Regulations
      • Attendance and Absences
      • Interruption of Degree Work
      • Graduation Eligibility
    • Degree and Diploma Programs
      • Bachelor of Fine Arts in Dance (BFA)
      • Bachelor of Music (BM)
        • Curricula
          • Bachelor of Music in Composition
          • Bachelor of Music in Jazz Performance
          • Bachelor of Music in Music Education
            • Bachelor of Music in Music Education -​ Composition
            • Bachelor of Music in Music Education-​ Guitar
            • Bachelor of Music in Music Education -​ Jazz
            • Bachelor of Music in Music Education -​ Orchestral Instruments
            • Bachelor of Music in Music Education -​ Piano
            • Bachelor of Music in Music Education -​ Voice
          • Bachelor of Music in Music for New Media
          • Bachelor of Music in Performance
            • Bachelor of Music in Performance -​ Computer Music
            • Bachelor of Music in Performance -​ Guitar
            • Bachelor of Music in Performance -​ Harpsichord
            • Bachelor of Music in Performance -​ Historical Performance
            • Bachelor of Music in Performance -​ Orchestral Instruments
            • Bachelor of Music in Performance -​ Organ
            • Bachelor of Music in Performance -​ Piano
            • Bachelor of Music in Performance -​ Voice
          • Bachelor of Music in Recording Arts &​ Sciences
            • Bachelor of Music in Recording Arts &​ Sciences -​ Composition
            • Bachelor of Music in Recording Arts &​ Sciences -​ Computer Music
            • Bachelor of Music in Recording Arts &​ Sciences -​ Guitar
            • Bachelor of Music in Recording Arts &​ Sciences -​ Jazz
            • Bachelor of Music in Recording Arts &​ Sciences -​ Orchestral Instruments
            • Bachelor of Music in Recording Arts &​ Sciences -​ Piano
        • Minors
          • Business of Music, Minor
          • Directed Studies, Minor
          • Historical Performance, Minor
          • Historical Performance: Voice, Minor
          • Liberal Arts, Minor
          • Music Theory, Minor
          • Musicology, Minor
        • Combined Degree Programs
          • Peabody-​Homewood Double Degree Program
        • Accelerated Graduate Degrees
          • Five-​Year BM/​MM Program
          • Five-​Year BMRA/​MA Program
            • Five-​Year BM/​MA: Music for New Media Variant
      • Master of Music (MM)
        • Master of Music in Composition
        • Master of Music in Film and Game Scoring
        • Master of Music: Performance
          • Master of Music, Performance -​ Choral Conducting Specialization
          • Master of Music, Performance -​ Computer Music specialization
          • Master of Music, Performance -​ Guitar specialization
          • Master of Music, Performance -​ Harpsichord specialization
          • Master of Music, Performance -​ Historical Performance Instruments specialization
          • Master of Music, Performance -​ Historical Performance Voice specialization
          • Master of Music, Performance -​ Jazz specialization
          • Master of Music, Performance -​ Orchestral Conducting specialization
          • Master of Music, Performance -​ Orchestral Instruments specialization
          • Master of Music, Performance -​ Organ specialization
          • Master of Music, Performance -​ Piano specialization
          • Master of Music, Performance -​ Wind Conducting specialization
          • Master of Music, Performance -​ Voice specialization
        • Master of Music: Academic Majors
          • Performance, Master of Music -​ Pedagogy emphasis
          • Music Education, Master of Music
          • Musicology, Master of Music
          • Music Theory Pedagogy, Master of Music
        • Master of Music: Low Residency
      • Master of Arts (MA)
        • Audio Sciences: Acoustics, Master of Arts
          • Five-​Year BM/​MA Program Requirements: Acoustics
        • Audio Sciences: Recording Arts and Sciences, Master of Arts
          • Five-​Year BM/​MA Program Requirements: Recording Arts
      • Doctor of Musical Arts (DMA)
        • Composition, Doctor of Musical Arts
        • Performance, Doctor of Musical Arts -​ Guitar specialization
        • Performance, Doctor of Musical Arts -​ Historical Performance Instruments specialization
        • Performance, Doctor of Musical Arts -​ Orchestral Conducting specialization
        • Performance, Doctor of Musical Arts -​ Orchestral Instruments specialization
        • Performance, Doctor of Musical Arts -​ Organ specialization
        • Performance, Doctor of Musical Arts -​ Piano specialization
        • Performance, Doctor of Musical Arts -​ Voice specialization
        • Performance, Doctor of Musical Arts -​ Wind Conducting specialization
      • Performer’s Certificate (PC)
        • Guitar, Performer's Certificate
        • Orchestral Instruments, Performer's Certificate
        • Organ, Performer's Certificate
        • Piano, Performer's Certificate
        • Voice, Performer's Certificate
      • Graduate Performance Diploma (GPD)
      • Artist’s Diploma (AD)
    • Extension Study
      • Music Education Certification -​ Instrumental
      • Music Education Certification -​ Vocal
  • Nitze School of Advanced International Studies
    • Degrees and Certificates
      • International Studies, Doctor of Philosophy
      • International Affairs, Doctor of
      • European Public Policy, Master of Arts
      • Global Policy, Master of Arts
      • Global Risk, Master of Arts (On-​site)
      • Global Risk, Master of Arts (Online)
      • International Affairs, Master of Arts
      • International Economics and Finance, Master of Arts
      • International Relations, Master of Arts
      • International Studies, Master of Arts
      • International Public Policy, Master of
      • Strategy, Cybersecurity, and Intelligence, Master of Arts
      • Sustainable Energy, Master of Arts (Online)
      • Chinese and American Studies, Hopkins-​Nanjing Center Certificate
      • Dual Degrees and Exchange Programs
      • Graduate Certificates
      • International Studies, Diploma
    • Policies and Resources
      • Academic Integrity
      • Academic Policies and Resources
      • Student Life
    • School Leadership and Key Contacts
  • School of Education
    • Academic and Student Policies
      • Academic and Student Conduct Policies
      • Academic Standards
      • Grading System and Academic Records
      • Grievances and Complaints
    • Admission
    • Graduation
    • Programs
      • Doctoral Programs
        • Education (Online), EdD
        • Education, PhD
      • Master's Programs
        • Counseling, Master of Science
        • Education, Master of Science
          • Education, Master of Science – Digital Age Learning and Educational Technology (Online)
          • Education, Master of Science -​ Educational Studies
          • Education, Master of Science -​ Gifted Education
          • Education, Master of Science -​ School Administration and Supervision
        • Education Policy, Master of Science
        • Health Professions (Online), Master of Education
        • Special Education, Master of Science
        • Teaching Professionals, Master of Education
      • Post Master's Certificates
        • Applied Behavior Analysis, Post–Master’s Certificate
        • Clinical Mental Health Counseling, Post–Master’s Certificate
        • Evidence-​Based Teaching in the Health Professions, Post–Master’s Certificate
      • Certificate of Advanced Graduate Study
        • Counseling, Certificate of Advanced Graduate Study
      • Graduate Certificates
        • Education of Students with Autism and Other Pervasive Developmental Disorders, Graduate Certificate
        • Educational Leadership for Independent Schools, Graduate Certificate
        • Gifted Education, Graduate Certificate
        • Leadership in Technology Integration (Online), Graduate Certificate
        • Mathematics/​STEM Instructional Leader (PreK-​6) (Online), Graduate Certificates
        • Mind, Brain and Teaching (Online), Graduate Certificate
        • School Administration and Supervision, Graduate Certificate
        • Urban Education, Graduate Certificate
    • Centers &​ Institutes
    • Scholarships
    • State Authorization of Distance Education (NC-​SARA)
  • School of Medicine
    • General Information
      • Conduct in Teacher/​Learner Relationships (Student Mistreatment Policy)
      • Lectureships and Visiting Professorships
      • Loan Funds
      • Medical Student Advising
      • Named Professorships
      • Office of Medical Student Affairs
      • Scholarships
      • Student Research Scholarships and Awards
      • Tuition
      • Tuition and Other Fees
      • Young Investigators’ Day
    • Policies
    • Graduate Programs
      • Anatomy Education, MS
      • Applied Health Sciences Informatics, MS
      • Biochemistry, Cellular and Molecular Biology, PhD
      • Biological Chemistry, PhD
      • Biomedical Engineering, PhD
      • Biophysics and Biophysical Chemistry, PhD/​Molecular Biophysics, PhD
      • Cellular and Molecular Medicine, PhD
      • Cellular and Molecular Physiology, PhD
      • Clinical Anaplastology, MS
      • Clinical Informatics, Post-​Baccalaureate Certificate
      • Cross-​Disciplinary Program in Biomedical Sciences, PhD
      • Functional Anatomy and Evolution, PhD
      • Health Sciences Informatics, PhD
      • Health Sciences Informatics–Research, MS
      • History of Medicine, MA (On-​site)
      • History of Medicine, MA (Online)
      • History of Medicine, PhD
      • History of Medicine, Post-​Baccalaureate Certificate (Online)
      • Human Genetics and Genomics, PhD
      • Immunology, PhD
      • Medical and Biological Illustration, MA
      • Medical Physics, MS
      • Neuroscience, PhD
      • Pathobiology, PhD
      • Pharmacology, PhD
    • Medical Program
      • Doctor of Medicine, MD
      • MD-​PhD, Combined Degree
      • Subject Areas
        • Anesthesiology and Critical Care Medicine
        • Biological Chemistry
        • Biomedical Engineering
        • Biophysics and Biophysical Chemistry
        • Cell Biology
        • Department of Genetic Medicine
        • Dermatology
        • Emergency Medicine
        • Epidemiology
        • Functional Anatomy and Evolution
        • Gynecology and Obstetrics
        • Health Sciences Informatics
        • History of Medicine
        • Medicine
        • Molecular and Comparative Pathobiology
        • Molecular Biology and Genetics
        • Multi-​Department Courses
        • Neurology
        • Neuroscience
        • Oncology
        • Ophthalmology
        • Pathology
        • Pediatrics
        • Pharmacology and Molecular Sciences
        • Physical Medicine and Rehabilitation
        • Physiology
        • Psychiatry and Behavioral Sciences
        • Public Health
        • Radiation Oncology and Molecular Radiation Sciences
        • Radiology and Radiological Science
        • Section of Surgical Sciences
    • Postdoctoral Fellows
  • School of Nursing
    • Admission
    • Advising
    • Certificates
      • Healthcare Organizational Leadership, Post-​Master’s Certificate
      • Nursing Education, Post-​Master's Certificate
      • Pediatric Acute Care Nurse Practitioner, Post-​Master's Certificate
      • Psychiatric Mental Health Nurse Practitioner, Post-​Master's Certificate
    • Doctoral Degrees
      • Doctor of Nursing Practice, Advanced Practice Track
        • Adult-​Gerontological Acute Care Nurse Practitioner, DNP Advanced Practice Track
        • Adult-​Gerontological Critical Care Clinical Nurse Specialist, DNP Advanced Practice Track
        • Adult-​Gerontological Health Clinical Nurse Specialist, DNP Advanced Practice Track
        • Adult-​Gerontological Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Family Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Nurse Anesthesia, DNP Advanced Practice Track
        • Pediatric Critical Care Clinical Nurse Specialist, DNP Advanced Practice Track
        • Pediatric Dual Primary/​Acute Care Nurse Practitioner, DNP Advanced Practice Track
        • Pediatric Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Psychiatric Mental Health Nurse Practitioner, DNP Advanced Practice Track
      • Doctor of Nursing Practice: Executive Track
      • Nursing, Doctor of Philosophy
      • Doctor of Nursing Practice (DNP): Advanced Practice Track/​Doctor of Philosophy in Nursing (PhD) Dual Degree
    • Dual Degrees
      • DNP Executive/​MBA Dual Degree
      • DNP Executive/​MPH Dual Degree
      • Healthcare Organizational Leadership, MSN/​MBA, Dual Degree
    • Financial Aid
    • Master's Degrees
      • Entry into Nursing, Master of Science in Nursing
      • Healthcare Organizational Leadership Track, Master of Science in Nursing
    • Online Prerequisites for Health Professions
    • Policies
      • Academic Integrity Policy
      • Academic Standards for Progression
      • Administrative Leave
      • Absence and Attendance Policy
      • Canvas and SON IT Help
      • Clinical Placements
      • Clinical Warnings
      • Complaint/​Grievance Policy
      • Compliance
      • Course Policies
      • Criminal Conduct/​Background Check Policies
      • Drug Testing Policy
      • Email Policy
      • Examination Policy
      • Grading Policy
      • Health Insurance for Students
      • Incomplete Coursework
      • Independent Study Policy
      • Leave of Absence
      • Letters of Recommendation
      • NCLEX
      • Non-​Degree-​Seeking Students
      • Notification of Missed Clinical Time
      • Pet Guidelines
      • Printing and Copying
      • Professional Attire Policy
      • Professional Ethics Policy
      • Registration Policies and Procedures
      • Religious Observance Attendance Policy
      • Social Media Guidelines
      • Student Code of Conduct
      • Technical Standards for Admission and Graduation
      • Transcripts and Enrollment Verifications
      • Transfer of Graduate Credit
      • Withdrawal Policy
    • Student Accounts
    • Tuition and Fees
  • Whiting School of Engineering
    • Full-​time, On-​campus Undergraduate and Graduate Programs (Homewood)
      • Zanvyl Krieger School of Arts and Sciences &​ Whiting School of Engineering Full-​Time, On-​Campus Undergraduate and Graduate Policies
      • Departments, Program Requirements, and Courses
        • Applied Mathematics and Statistics
          • Applied Mathematics and Statistics, Bachelor of Arts
          • Applied Mathematics and Statistics, Bachelor of Science
          • Applied Mathematics and Statistics, Master of Science in Engineering
          • Applied Mathematics and Statistics, Minor
          • Applied Mathematics and Statistics, PhD
          • Data Science, Master's Degree
          • Financial Mathematics, Master of Science in Engineering
        • Biomedical Engineering
          • Bioengineering Innovation and Design, Master of Science in Engineering
          • Biomedical Engineering, Bachelor of Arts
          • Biomedical Engineering, Bachelor of Science
          • Biomedical Engineering, Master of Science in Engineering
          • Biomedical Engineering, PhD through the School of Medicine
        • Center for Leadership Education
          • Accounting and Financial Management, Minor
          • Engineering Management, Master of Science
          • Global Innovation and Leadership Through Engineering, Master of Science
          • Leadership Studies, Minor
          • Marketing and Communications, Minor
          • Professional Communication Program
          • Professional Development Program
          • W.P. Carey Entrepreneurship and Management, Minor
        • Chemical and Biomolecular Engineering
          • Chemical and Biomolecular Engineering, Bachelor of Science
          • Chemical and Biomolecular Engineering, Master of Science in Engineering
          • Chemical and Biomolecular Engineering, PhD
        • Civil &​ Systems Engineering
          • Civil Engineering, Bachelor of Science
          • Civil Engineering, Master of Science in Engineering (MSE)
          • Civil Engineering, Minor
          • Civil and Systems Engineering, PhD
          • Systems Engineering, Bachelor of Science
          • Systems Engineering, Master of Science
          • Systems Engineering, Minor
        • Computational Medicine
          • Computational Medicine, Minor
        • Computer Science
          • Computer Science, Bachelor of Arts
          • Computer Science, Bachelor of Science
          • Computer Science, Master of Science in Engineering
          • Computer Science, Minor
          • Computer Science, PhD
        • Doctor of Engineering
          • Engineering, Doctor of Engineering
        • Electrical and Computer Engineering
          • Computer Engineering, Bachelor of Science
          • Electrical and Computer Engineering, Master of Science in Engineering
          • Electrical and Computer Engineering, PhD
          • Electrical Engineering, Bachelor of Science
          • Energy, Minor
        • Environmental Health and Engineering
          • Engineering for Sustainable Development, Minor
          • Environmental Engineering, Bachelor of Science
          • Environmental Engineering, Minor
          • Environmental Sciences, Minor
          • Geography and Environmental Engineering, Master of Arts
          • Geography and Environmental Engineering, Master of Science
          • Geography and Environmental Engineering, Master of Science in Engineering
          • Geography and Environmental Engineering, PhD
          • Occupational and Environmental Hygiene, Master of Science
        • General Engineering
          • General Engineering, Bachelor of Arts
        • Information Security Institute
          • Security Informatics, Master of Science
          • Security Informatics, Master of Science/​Applied Mathematics and Statistics, Master of Science in Engineering Dual Master's Program
          • Security Informatics, Master of Science/​Computer Science, Master of Science in Engineering Dual Master's Program
        • Materials Science and Engineering
          • Materials Science and Engineering, Bachelor of Science
          • Materials Science and Engineering, Master of Science in Engineering
          • Materials Science and Engineering, PhD
        • Mechanical Engineering
          • Engineering Mechanics, Bachelor of Science
          • Mechanical Engineering, Bachelor of Science
          • Mechanical Engineering, Master of Science in Engineering
          • Mechanical Engineering, PhD
        • NanoBioTechnology
        • Robotics and Computational Sensing
          • Computer Integrated Surgery, Minor
          • Robotics, Master of Science in Engineering
          • Robotics, Minor
      • Multi-​School Programs of Study
        • Business, Minor
        • Peabody-​Homewood Double Degree Program
        • Space Science and Engineering
    • Part-​Time, Online Graduate Programs (Engineering for Professionals)
      • Academic Policies
        • Academic Calendar
        • Academic Regulations
        • Registration Policies
        • Tuition and Fees
      • Admission Requirements
      • Applied and Computational Mathematics
        • Applied and Computational Mathematics, Graduate Certificate
        • Applied and Computational Mathematics, Master of Science
        • Applied and Computational Mathematics, Post-​Master’s Certificate
      • Applied Biomedical Engineering
        • Applied Biomedical Engineering, Graduate Certificate
        • Applied Biomedical Engineering, Master of Science
        • Applied Biomedical Engineering, Post-​Master’s Certificate
      • Applied Physics
        • Applied Physics, Master of Science
        • Applied Physics, Post-​Master’s Certificate
      • Artificial Intelligence
        • Artificial Intelligence, Graduate Certificate
        • Artificial Intelligence, Master of Science
      • Chemical and Biomolecular Engineering
        • Chemical and Biomolecular Engineering, Master of Chemical and Biomolecular Engineering
      • Civil Engineering
        • Civil Engineering, Graduate Certificate
        • Civil Engineering, Master of Civil Engineering
      • Computer Science
        • Computer Science, Graduate Certificate
        • Computer Science, Master of Science
        • Computer Science, Post-​Master’s Certificate
      • Cybersecurity
        • Cybersecurity, Graduate Certificate
        • Cybersecurity, Master of Science
        • Cybersecurity, Post-​Master’s Certificate
      • Data Science
        • Data Science, Graduate Certificate
        • Data Science, Master of Science
        • Data Science, Post-​Master’s Certificate
      • Electrical and Computer Engineering
        • Electrical and Computer Engineering, Graduate Certificate
        • Electrical and Computer Engineering, Master of Science
        • Electrical and Computer Engineering, Post-​Master’s Certificate
      • Engineering Management
        • Engineering Management, Graduate Certificate
        • Engineering Management, Master of Engineering Management
      • Environmental Engineering, Science, Management, and Sustainability Programs
        • Climate Change, Energy, and Environmental Sustainability, Graduate Certificate
        • Climate, Energy, and Environmental Sustainability, Master of Science
        • Environmental Engineering
          • Environmental Engineering, Graduate Certificate
          • Environmental Engineering, Master of Environmental Engineering
          • Environmental Engineering, Post-​Master’s Certificate
        • Environmental Engineering and Science
          • Environmental Engineering and Science, Graduate Certificate
          • Environmental Engineering and Science, Master of Science
          • Environmental Engineering and Science, Post-​Master’s Certificate
        • Environmental Planning and Management
          • Environmental Planning and Management, Graduate Certificate
          • Environmental Planning and Management, Master of Science
          • Environmental Planning and Management, Post-​Master’s Certificate
      • Financial Mathematics
        • Financial Mathematics, Master of Science
        • Financial Risk Management, Graduate Certificate
        • Quantitative Portfolio Management, Graduate Certificate
        • Securitization, Graduate Certificate
      • Healthcare Systems Engineering
        • Healthcare Systems Engineering, Master of Science
      • Industrial and Operations Engineering
        • Industrial and Operations Engineering, Master of Science
      • Information Systems Engineering
        • Information Systems Engineering, Graduate Certificate
        • Information Systems Engineering, Master of Science
        • Information Systems Engineering, Post-​Master’s Certificate
      • Materials Science and Engineering
        • Materials Science and Engineering, Master of Science
      • Mechanical Engineering
        • Mechanical Engineering, Master of Science
        • Mechanical Engineering, Post-​Master’s Certificate
      • Occupational and Environmental Hygiene
        • Occupational and Environmental Hygiene, Master of Science
      • Robotics and Autonomous Systems
        • Robotics and Autonomous Systems, Master of Science
      • Space Systems Engineering
        • Space Systems Engineering, Master of Science
        • Space Systems Engineering, Post-​Master's Certificate
      • Systems Engineering
        • Systems Engineering, Graduate Certificate
        • Systems Engineering, Master of Science
        • Systems Engineering, Master of Science in Engineering (ABET-​accredited)
        • Systems Engineering, Post-​Master’s Certificate
  • Zanvyl Krieger School of Arts and Sciences
    • Full-​time, On-​campus Undergraduate and Graduate Programs (Homewood)
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  • Home›
  • School of Medicine›
  • Graduate Programs›
  • Neuroscience, PhD
  • Overview
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The Department of Neuroscience offers an interdisciplinary program designed to train doctoral students for independent research and teaching in neuroscience. It is the goal of the program to ensure that candidates for the Ph.D. and M.D./Ph.D. degrees obtain a background covering molecular, cellular, systems, and cognitive approaches to neuroscience, as well as receive training that brings them to the forefront of research in their particular area of interest. A series of core courses in neuroscience, along with advanced electives, seminar series, laboratory rotations, and original independent dissertation research, form the Neuroscience Graduate Training Program.

Students enter the program from different backgrounds and the laboratories in which they elect to work cover different disciplines; therefore, the program is tailored to fit the needs of individual students. The academic year at the Johns Hopkins University School of Medicine is divided into four quarters plus a summer semester. Courses are designed so that students have ample time to become involved in laboratory rotations. These laboratory rotations expose the student to a variety of current research techniques in neuroscience and provide an opportunity for the student to select a laboratory in which to conduct dissertation research. Scheduling of the three rotations is adjusted to make the most convenient schedule for each student. The rotations are usually completed by the end of the first full year in the program. Most students begin their thesis research at the beginning of their second year.

For more information, please visit The Solomon H. Snyder Department of Neuroscience webpage: http://neuroscience.jhu.edu.

Financial Aid

The program provides tuition remission plus a stipend at or above the National Institutes of Health Predoctoral level for all students. All entering and first-year students are encouraged to apply for individual fellowships such as those sponsored by the National Science Foundation and the Howard Hughes Medical Institute.

Vivien Thomas PhD Scholars at JHU
The Vivien Thomas Scholars Initiative (VTSI) is a new endowed fellowship program at Johns Hopkins for PhD students in STEM fields. It provides full tuition, stipend, and benefits while also providing targeted mentoring, networking, community, and professional development opportunities. Students who have attended a historically black college and university (HBCU) or other minority serving institution (MSI) for undergraduate study are eligible to apply. More information about the VTSI program is available at this link: https://provost.jhu.edu/about/vivien-thomas-scholars-initiative/. To be considered for the VTSI, all application and supplementary materials must be received by December 1st.

Admission Requirements

We use a holistic approach to evaluating applicants and look forward to reading your application. We are most enthusiastic about applicants who have taken full advantage of the opportunities available at their undergraduate institution and through other summer or postbac experiences. Our class size is typically ~18 students per year.

Applicants are expected to have received a B.S. or B.A. prior to enrolling in the graduate program. Laboratory research experience prior to enrollment is also desirable. If you have research experience, please describe your research in your Statement of Interest and Career Objectives and indicate the number of months engaged in full-time and part-time research on your CV. Students who do well in our program typically have a strong academic foundation in areas of biological or physical sciences. Some of the courses that prepare students well include general biology, neuroscience, mathematics through calculus, general physics, general chemistry, organic chemistry, statistics, engineering, or computer science.

NOTE: The Neuroscience Program DOES NOT require GRE scores. 

Program Requirements

Courses

A year-long core course provides an integrated overview of molecular and cellular neuroscience, neuroanatomy and systems, and cognitive neuroscience. This course is aimed at providing Neuroscience graduate students with a foundation for posing meaningful questions in their area of interest.  During the first two years, students are required to take 6 graduate level core courses that provide rigorous training in principles of neuroscience research. In addition, students in the first year attend research symposia and complete lab rotations to introduce them to research. Students in the program are also required to participate in core program activities such as seminars, journal clubs, a quantitative analysis boot camp, career development courses and various program events. In addition, each student selects advanced electives offered by members of the Neuroscience Training Program or other departments at the Medical School.

Seminar Program

The Neuroscience Training Program conducts several seminar series to ensure that students are exposed to recent work by researchers from across the country and the world as well as by Hopkins faculty and fellows. Graduate trainees participate actively in these series throughout their training, including inviting and hosting three speakers each year. A weekly lecture is given by an outstanding researcher in some field of neuroscience. Seminars are selected so that an overall balance of subject matter is covered yearly. Students are given an opportunity to meet with each speaker for questions and discussion. Weekly lunchtime talks are presented on current literature by graduate students and postdoctoral fellows. Since an ability to communicate scientific work clearly is essential, graduate students receive close guidance in preparing and evaluating their journal club presentations. Once a month, the faculty, postdoctoral fellows, and students from one laboratory present and discuss the ongoing research in that laboratory. This provides an informal setting to discuss research being conducted in the laboratories of the Neuroscience Training Program and gives advanced graduate students and postdoctoral fellows a forum for presenting their work.

Requirements for the PhD Degree

A minimum residency of two academic years is required. During the course of graduate study, the student must successfully complete the required course requirements. An oral examination, conducted as prescribed by the Doctor of Philosophy Board, must be completed by the end of the second year. The student must then conduct original research and describe this research in a written thesis dissertation, which must be approved by the students Thesis Committee and the Doctor of Philosophy Board.

Training Facilities

The Training Program is centered in the Department of Neuroscience. The Training Program utilizes laboratory facilities located in the Department of Neuroscience plus several other basic and clinical departments closely associated with the Neuroscience Department. All of these laboratories are within a short distance of each other. Modern state of the art facilities for research in molecular biology, neurophysiology, pharmacology, biochemistry, cell biology, and morphology are available. The Mind/Brain Institute, located on the Homewood Campus of the University, is a group of laboratories devoted to the investigation of the neural mechanisms of higher mental function and particularly to the mechanisms of perception. All of the disciplines required to address these questions are represented in the Institute. These include neurophysiology, psychology, theoretical neurobiology, neuroanatomy, and cognitive science. All of the faculty in the Mind/Brain Institute are members of the Neuroscience Graduate Program.

Combined M.D./Ph.D. Program

A subset of the current predoctoral trainees in the Neuroscience Program are candidates for both Ph.D. and M.D. degrees. Applications for admission to the combined program are considered by the M.D./Ph.D. Committee of the School of Medicine. Application forms for the School of Medicine contain a section requesting information relevant to graduate study. Applicants interested in the combined M.D./Ph.D. program should complete this section also, and indicate specifically their interest in the “Neuroscience Training Program”. If application to the combined M.D./Ph.D. program proves unsuccessful and the applicant wishes to be considered for graduate studies, they must notify the Admissions Office of the Neuroscience Training Program by separate letter.

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