• Skip to Content
  • AZ Index
  • Catalogue Home
  • Johns Hopkins University Home
Johns Hopkins University
Academic Catalogue | 2022-23 Edition
Class Schedule Search
Search location
  • Catalogue Home
  • Programs
  • Courses
  • Policies & Information
  • Print Options
  • Archives
  • Amendments

Immunology, PhD

School of Medicine

Catalogue Home

  • Explore our Programs
  • University-​wide Policies and Information
    • Academic Policies and Information
      • Academic Calendar
      • Academic Integrity Policies
      • Animal Care and Use Program
      • Credit Hour Policy
      • FERPA
      • PHD Specific Policies
      • Transcripts and Enrollment Verifications
    • Admission and Aid
      • Tuition and Fees
        • Financial Aid
    • Higher Education Act Disclosures
      • General Institutional Information
      • Health and Safety Information
      • Student Financial Assistance Information
    • Office of Institutional Equity
      • Discrimination and Harassment Policy and Procedures
      • Equal Opportunity and Title IX Notice
      • Sexual Misconduct
    • Rights, Privileges, and Responsibilities
      • Academic Grievance Policy: Students and Postdoctoral Fellows
      • New Child Accommodations for Full-​Time Graduate Students and Postdoctoral Trainees
      • Personal Relationships Policy
      • Photography and Film Rights Policy
      • Student Conduct Code
      • Student Disability Services (SDS)
      • Student Health
    • Veterans Affairs
  • Bloomberg School of Public Health
    • Academic Calendar
    • Admission
    • CEPH Requirements
    • Departments
      • Department of Biochemistry and Molecular Biology
        • Biochemistry and Molecular Biology, MHS
        • Biochemistry and Molecular Biology, ScM
        • Biochemistry and Molecular Biology, PhD
        • Non-​Degree Training
      • Department of Biostatistics
        • Biostatistics, MHS
        • Biostatistics, ScM
        • Biostatistics, PhD
      • Department of Environmental Health and Engineering
        • Environmental Health, MHS
        • Environmental Health, SCM
        • Toxicology for Human Risk Assessment, MS
        • Environmental Health, PhD
        • Non-​Degree Training
      • Department of Epidemiology
        • Epidemiology, MHS
        • Epidemiology, ScM
        • Epidemiology, PhD
        • Non-​Degree Training
      • Department of Health, Behavior and Society
        • Social Factors in Health, MHS
        • Health Education and Health Communication, MSPH
        • Genetic Counseling, ScM
        • Health, Behavior and Society, PhD
        • Non-​Degree Training
      • Department of Health Policy and Management
        • Health Administration, MHA
        • Health Policy, MSPH
        • Health Economics and Outcomes Research, MHS
        • Health Policy and Management, PhD
        • Health Policy and Management, DrPH (Tsinghua)
        • Non-​Degree Training
      • Department of International Health
        • Global Health Economics, MHS
        • International Health, MSPH
        • International Health, MSPH, Human Nutrition-​Dietitian
        • International Health, MA/​MSPH
        • International Health, PhD
        • Non-​Degree Training
      • Department of Mental Health
        • Mental Health, MHS
        • Mental Health, PhD
        • Non-​Degree Training
      • Department of Molecular Microbiology &​ Immunology
        • Molecular Microbiology &​ Immunology, MHS
        • Molecular Microbiology &​ Immunology, ScM
        • Molecular Microbiology &​ Immunology, PhD
        • Non-​Degree Training
      • Department of Population, Family and Reproductive Health
        • Population, Family and Reproductive Health, MHS
        • Population, Family and Reproductive Health, MHS Online
        • Population, Family and Reproductive Health, MSPH
        • Population, Family and Reproductive Health, PhD
      • Doctor of Public Health (DrPH)
      • Graduate Training Programs in Clinical Investigation
        • Graduate Training Programs in Clinical Investigation, MHS
        • Graduate Training Programs in Clinical Investigation, PhD
      • Master of Arts in Public Health Biology
      • Master of Bioethics
      • Master of Public Health Program
        • DNP/​MPH
        • DVM/​MPH
        • JD/​MPH
        • LLM/​MPH
        • MBA/​MPH with China Europe International Business School
        • MD/​MPH
        • MPH/​MBA
        • MSW/​MPH
      • Online Programs for Applied Learning (OPAL)
        • Master of Applied Science in Community-​Based Primary Health Care Programs in Global Health
        • Master of Applied Science in Global Health Planning and Management
        • Master of Applied Science in Humanitarian Health
        • Master of Applied Science in Patient Safety and Healthcare Quality
        • Master of Applied Science in Population Health Management
        • Master of Applied Science in Spatial Analysis for Public Health
      • Residency Programs
        • General Preventive Medicine Residency Program
        • Occupational and Environmental Medicine Residency
    • Certificates
      • Adolescent Health, Certificate
      • Bioethics, Certificate
      • Climate and Health, Certificate
      • Clinical Trials, Certificate
      • Community-​Based Public Health, Certificate
      • Demographic Methods, Certificate
      • Environmental and Occupational Health, Certificate
      • Epidemiology for Public Health Professionals, Certificate
      • Evaluation: International Health Programs, Certificate
      • Food Systems, the Environment &​ Public Health, Certificate
      • Gender and Health, Certificate
      • Gerontology, Certificate
      • Global Health, Certificate
      • Global Health Practice, Certificate
      • Global Tobacco Control, Certificate
      • Health and Human Rights, Certificate
      • Health Communication, Certificate
      • Health Disparities and Health Inequality, Certificate
      • Health Education, Certificate
      • Health Finance and Management, Certificate
      • Healthcare Epidemiology and Infection Prevention and Control, Certificate
      • Humane Sciences and Toxicology Policy, Certificate
      • Humanitarian Health, Certificate
      • Injury and Violence Prevention, Certificate
      • International Healthcare Management and Leadership, Certificate
      • Leadership for Public Health and Healthcare, Certificate
      • Lesbian, Gay, Bisexual, Transgender, and Queer (LGBTQ) Public Health, Certificate
      • Maternal and Child Health, Certificate
      • Mental Health Policy, Economics and Services, Certificate
      • Pharmacoepidemiology and Drug Safety, Certificate
      • Population and Health, Certificate
      • Population Health Management, Certificate
      • Product Stewardship for Sustainability, Certificate
      • Public Health Advocacy, Certificate
      • Public Health Economics, Certificate
      • Public Health Informatics, Certificate
      • Public Health Practice, Certificate
      • Public Health Preparedness, Certificate
      • Public Health Training Certificate for American Indian Health Professionals
      • Public Mental Health Research, Certificate
      • Quality, Patient Safety, and Outcomes Research, Certificate
      • Quantitative Methods in Public Health, Certificate
      • Rigor, Reproducibility and Responsibility in Scientific Practice, Certificate
      • Risk Sciences and Public Policy, Certificate
      • Spatial Analysis for Public Health, Certificate
      • Training Certificate in Public Health
      • Tropical Medicine, Certificate
      • Vaccine Science and Policy, Certificate
    • Policies
      • Academic
        • Academic Ethics Code
        • Academic Leave of Absence
        • Compliance Line
        • Grade Appeal Policy
        • Grading System
        • Graduation Policy
        • Interdivisional Registration
        • Involuntary Leave of Absence
        • Multi-​Term Course Policy
        • Post-​Doctoral Fellow Student Status
        • Student Grievance Policy
      • Research
        • Animal Research
        • Human Subjects Research
        • Worker's Comp
      • Student Life
        • Alternative Beverages
        • Donation Drive Protocol
        • Social Media Policy
        • Special Events Coordination
        • Student Fundraising
  • Carey Business School
    • Admission
      • Graduate Degree Requirements
      • Master’s Programs
      • Certificate Programs
      • Verification of Credentials
      • International Student Admission Policy
      • Inactive/​Deactivated Certificate or Degree Applications
      • State-​Specific Authorization for Online Courses
    • Degrees and Certificates
      • Business Administration (Flexible), MBA
      • Business Administration (Full Time), MBA
      • Business Analytics and Risk Management (Part Time), Master of Science
      • Business Analytics and Risk Management, Master of Science
      • Design Leadership, MBA/​MA Dual Degree
      • Finance (Part Time), Master of Science
      • Finance, Master of Science
      • Financial Management, Graduate Certificate
      • Financial Management, Graduate Certificate, Investments, Graduate Certificate, Applied Economics, MS
      • Health Care Management (Part Time), Master of Science
      • Health Care Management, Master of Science
      • Information Systems, Master of Science
      • Investments, Graduate Certificate
      • Leadership Development Program, Graduate Certificate
      • Marketing (Part Time), Master of Science
      • Marketing, Master of Science
      • MBA/​Applied Economics, MS Dual Degree
      • MBA/​Biotechnology, MS Dual Degree
      • MBA/​Communication, MA Dual Degree
      • MBA/​DNP Dual Degree
      • MBA/​Government, MA Dual Degree
      • MBA/​Healthcare Organizational Leadership, MSN Dual Degree
      • MBA/​JD Dual Degree
      • MBA/​MA in International Relations
      • MBA/​MD Dual Degree
      • MBA/​MPH Dual Degree
      • MSF/​MBA Dual Degree
      • Real Estate and Infrastructure (Part Time), Master of Science
      • Real Estate and Infrastructure, Master of Science
      • Business, Minor
    • Policies and Resources
      • Academic Ethics Policy
      • Academic Progress and Standards
      • Changing Degree Program
      • Grading Policy
      • Graduation
      • Attendance Policy
      • Leave of Absence
      • Registration
      • Student Accounts
      • Transfer of Graduate Credit
      • Waiver Exams
  • Peabody Institute
    • General Information, Procedures and Regulations
      • Introduction and Nomenclature
      • Mission
      • Accreditation
      • Links
      • Honor Societies
    • Procedural Information
      • Applicability
      • Studio Assignments
      • Course Numbering
      • Large Ensemble Participation
      • Competitions
      • Recitals
      • Academic Advising
      • Inter-​Institutional Academic Arrangements
      • Study Abroad Program
      • Outside Instruction and Public Performance
    • Academic Regulations
      • Applicability
      • Academic Code of Conduct
      • Program Classification, Status, and Credit Limits
      • Sources of Credit
      • Grading System and Regulations
      • Dean's List Criteria
      • Academic Standing
      • Registration Regulations
      • Attendance and Absences
      • Interruption of Degree Work
      • Graduation Eligibility
    • Degree and Diploma Programs
      • Bachelor of Music (BM)
        • Curricula
          • Bachelor of Music in Performance
            • Composition, Bachelor of Music
            • Computer Music, Bachelor of Music
            • Guitar, Bachelor of Music
            • Harpsichord, Bachelor of Music
            • Historical Performance, Bachelor of Music
            • Jazz, Bachelor of Music
            • Music for New Media, Bachelor of Music
            • Orchestral Instruments, Bachelor of Music
            • Organ, Bachelor of Music
            • Piano, Bachelor of Music
            • Voice, Bachelor of Music
          • Bachelor of Music in Music Education
            • Composition, Bachelor of Music Education
            • Guitar, Bachelor of Music Education
            • Jazz, Bachelor of Music Education
            • Orchestral Instruments, Bachelor of Music Education
            • Piano, Bachelor of Music Education
            • Voice, Bachelor of Music Education
          • Bachelor of Music in Recording Arts
            • Composition, Bachelor of Music in Recording Arts
            • Computer Music, Bachelor of Music in Recording Arts
            • Guitar, Bachelor of Music in Recording Arts
            • Jazz, Bachelor of Music in Recording Arts
            • Orchestral Instruments, Bachelor of Music in Recording Arts
            • Piano, Bachelor of Music in Recording Arts
        • Minors
          • Business of Music, Minor
          • Directed Studies, Minor
          • Historical Performance, Minor
          • Historical Performance: Voice, Minor
          • Liberal Arts, Minor
          • Music Theory, Minor
          • Musicology, Minor
        • Combined Degree Programs
          • Peabody-​Homewood Double Degree Program
        • Accelerated Graduate Degrees
          • Five-​Year BM/​MM Program
          • Five-​Year BMRA/​MA Program
            • Five-​Year BM/​MA: Music for New Media Variant
      • Bachelor of Fine Arts (BFA)
        • Minors
      • Master of Music (MM)
        • Master of Music: Performance
          • Composition, Master of Music
          • Computer Music, Master of Music
          • Guitar, Master of Music
          • Harpsichord, Master of Music
          • Historical Performance Instruments, Master of Music
          • Historical Performance Voice, Master of Music
          • Jazz, Master of Music
          • Orchestral Conducting, Master of Music
          • Orchestral Instruments, Master of Music
          • Organ, Master of Music
          • Piano, Master of Music
          • Piano: Ensemble Arts Vocal Accompanying, Master of Music
          • Wind Conducting, Master of Music
          • Voice, Master of Music
        • Master of Music: Academic Majors
          • Performance/​Pedagogy, Master of Music
          • Music Education, Master of Music
          • Musicology, Master of Music
          • Music Theory Pedagogy, Master of Music
        • Master of Music: Low Residency
      • Master of Arts (MA)
        • Acoustics, Master of Arts
          • Five-​Year BM/​MA Program Requirements: Acoustics
        • Recording Arts and Sciences, Master of Arts
          • Five-​Year BM/​MA Program Requirements: Recording Arts
      • Doctor of Musical Arts (DMA)
        • Composition, Doctor of Musical Arts
        • Guitar, Doctor of Musical Arts
        • Historical Performance Instruments, Doctor of Musical Arts
        • Orchestral Conducting, Doctor of Musical Arts
        • Orchestral Instruments, Doctor of Musical Arts
        • Organ, Doctor of Musical Arts
        • Piano, Doctor of Musical Arts
        • Voice, Doctor of Musical Arts
        • Wind Conducting, Doctor of Musical Arts
      • Performer’s Certificate (PC)
        • Guitar, Performer's Certificate
        • Orchestral Instruments, Performer's Certificate
        • Organ, Performer's Certificate
        • Piano, Performer's Certificate
        • Voice, Performer's Certificate
      • Graduate Performance Diploma (GPD)
      • Artist’s Diploma (AD)
    • Extension Study
      • Music Education Certification -​ Instrumental
      • Music Education Certification -​ Vocal
  • Nitze School of Advanced International Studies
    • Academic Policies and Resources
    • Degrees and Certificates
      • International Studies, Doctor of Philosophy
      • International Affairs, Doctor of
      • European Public Policy, Master of Arts
      • Global Policy, Master of Arts
      • Global Risk, Master of Arts (On-​site)
      • Global Risk, Master of Arts (Online)
      • International Affairs, Master of Arts
      • International Economics and Finance, Master of Arts
      • International Relations, Master of Arts
      • International Studies, Master of Arts
      • International Public Policy, Master of
      • Strategy, Cybersecurity, and Intelligence, Master of Arts
      • Sustainable Energy, Master of Arts (Online)
      • Chinese and American Studies, Hopkins-​Nanjing Center Certificate
      • Dual Degrees and Exchange Programs
      • Graduate Certificates
      • International Studies, Diploma
  • School of Education
    • Academic and Student Policies
      • Academic and Student Conduct Policies
      • Academic Standards
      • Grading System and Academic Records
      • Grievances and Complaints
    • Admission
    • Graduation
    • Programs
      • Doctoral Programs
        • Education (Online), EdD
        • Education, PhD
      • Master's Programs
        • Counseling, Master of Science
        • Education, Master of Science
        • Health Professions (Online), Master of Education
        • Special Education, Master of Science
      • Post Master's Certificates
        • Applied Behavior Analysis, Post–Master’s Certificate
        • Clinical Mental Health Counseling, Post–Master’s Certificate
        • Evidence-​Based Teaching in the Health Professions, Post–Master’s Certificate
      • Certificate of Advanced Graduate Study
        • Counseling, Certificate of Advanced Graduate Study
      • Graduate Certificates
        • Education of Students with Autism and Other Pervasive Developmental Disorders, Graduate Certificate
        • Educational Leadership for Independent Schools, Graduate Certificate
        • Gifted Education, Graduate Certificate
        • Leadership in Technology Integration (Online), Graduate Certificate
        • Mathematics/​STEM Instructional Leader (PreK-​6) (Online), Graduate Certificate
        • Mind, Brain and Teaching (Online), Graduate Certificate
        • School Administration and Supervision, Graduate Certificate
        • Urban Education, Graduate Certificate
    • Research and Development Centers
    • Scholarships
    • State Authorization of Distance Education and Higher Education Agencies in Other States
  • School of Medicine
    • General Information
      • Conduct in Teacher/​Learner Relationships (Student Mistreatment Policy)
      • Faculty Traveling Fellowship and Visiting Scholar Fellowship
      • Lectureships and Visiting Professorships
      • Loan Funds
      • Medical Student Advising
      • Named Professorships
      • Office of Medical Student Affairs
      • Scholarships
      • Student Research Scholarships and Awards
      • Tuition
      • Tuition and Other Fees
      • Young Investigators’ Day
    • Policies
    • Graduate Programs
      • Anatomy Education, MS
      • Applied Health Sciences Informatics, MS
      • Biochemistry, Cellular and Molecular Biology, PhD
      • Biological Chemistry, PhD
      • Biomedical Engineering, PhD
      • Biophysics and Biophysical Chemistry, PhD/​Molecular Biophysics, PhD
      • Cellular and Molecular Medicine, PhD
      • Cellular and Molecular Physiology, PhD
      • Clinical Anaplastology, MS
      • Clinical Informatics, Post-​Baccalaureate Certificate
      • Cross-​Disciplinary Program in Biomedical Sciences, PhD
      • Functional Anatomy and Evolution, PhD
      • Health Sciences Informatics, PhD
      • Health Sciences Informatics–Research, MS
      • History of Medicine, MA (On-​site)
      • History of Medicine, MA (Online)
      • History of Medicine, PhD
      • History of Medicine, Post-​Baccalaureate Certificate (Online)
      • Human Genetics and Molecular Biology, PhD
      • Immunology, PhD
      • Medical and Biological Illustration, MA
      • Medical Physics, MS
      • Neuroscience, PhD
      • Pathobiology, PhD
      • Pharmacology, PhD
    • Medical Program
      • Doctor of Medicine, MD
      • MD-​PhD, Combined Degree
      • Subject Areas
        • Anesthesiology and Critical Care Medicine
        • Biological Chemistry
        • Biomedical Engineering
        • Biophysics and Biophysical Chemistry
        • Cell Biology
        • Dermatology
        • Emergency Medicine
        • Epidemiology
        • Functional Anatomy and Evolution
        • Gynecology and Obstetrics
        • Health Sciences Informatics
        • History of Medicine
        • Institute of Genetic Medicine
        • Medicine
        • Molecular and Comparative Pathobiology
        • Molecular Biology and Genetics
        • Multi-​Department Courses
        • Neurology
        • Neuroscience
        • Oncology
        • Ophthalmology
        • Pathology
        • Pediatrics
        • Pharmacology and Molecular Sciences
        • Physical Medicine and Rehabilitation
        • Physiology
        • Psychiatry and Behavioral Sciences
        • Public Health
        • Radiation Oncology and Molecular Radiation Sciences
        • Radiology and Radiological Science
        • Section of Surgical Sciences
    • Postdoctoral Fellows
  • School of Nursing
    • Admission
    • Advising
    • Certificates
      • Healthcare Organizational Leadership, Post-​Master’s Certificate
      • Nursing Education, Post-​Master's Certificate
      • Pediatric Acute Care Nurse Practitioner, Post-​Master's Certificate
      • Psychiatric Mental Health Nurse Practitioner, Post-​Master's Certificate
    • Doctoral Degrees
      • Doctor of Nursing Practice, Advanced Practice Track
        • Adult-​Gerontological Acute Care Nurse Practitioner, DNP Advanced Practice Track
        • Adult-​Gerontological Critical Care Clinical Nurse Specialist, DNP Advanced Practice Track
        • Adult-​Gerontological Health Clinical Nurse Specialist, DNP Advanced Practice Track
        • Adult-​Gerontological Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Family Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Nurse Anesthesia, DNP Advanced Practice Track
        • Pediatric Critical Care Clinical Nurse Specialist, DNP Advanced Practice Track
        • Pediatric Dual Primary/​Acute Care Nurse Practitioner, DNP Advanced Practice Track
        • Pediatric Primary Care Nurse Practitioner, DNP Advanced Practice Track
        • Psychiatric Mental Health Nurse Practitioner, DNP Advanced Practice Track
      • Doctor of Nursing Practice: Executive Track
      • Nursing, Doctor of Philosophy
      • Doctor of Nursing Practice (DNP): Advanced Practice Track/​Doctor of Philosophy in Nursing (PhD) Dual Degree
    • Dual Degrees
      • DNP Executive/​MBA Dual Degree
      • DNP Executive/​MPH Dual Degree
      • Healthcare Organizational Leadership, MSN/​MBA, Dual Degree
    • Financial Aid
    • Master's Degrees
      • Entry into Nursing, Master of Science in Nursing
      • Healthcare Organizational Leadership Track, Master of Science in Nursing
    • Online Prerequisites for Health Professions
    • Policies
      • Academic Integrity Policy
      • Academic Standards for Progression
      • Administrative Leave
      • Attendance Policy
      • Canvas and SON IT Help
      • Clinical Placements
      • Clinical Warnings
      • Complaint/​Grievance Policy
      • Compliance
      • Continuous Enrollment Policy
      • Course Policies
      • Criminal Conduct Policy
      • Examination Policy
      • Grading Policy
      • Health Insurance for Students
      • Incomplete Coursework
      • Independent Study Policy
      • Involuntary Leave of Absence
      • Leave of Absence or Withdrawal
      • Letters of Recommendation
      • NCLEX
      • Non-​Degree-​Seeking Students
      • Notification of Missed Clinical Time
      • Pet Guidelines
      • Printing and Copying
      • Professional Attire Policy
      • Professional Ethics Policy
      • Registration Policies and Procedures
      • Religious Observance Attendance Policy
      • Student Code of Conduct
      • Technical Standards for Admission and Graduation
      • Transcripts and Enrollment Verifications
      • Transfer of Graduate Credit
    • Tuition and Fees
  • Whiting School of Engineering
    • Full-​time, On-​campus Undergraduate and Graduate Programs (Homewood)
      • Undergraduate Policies
        • Academic Policies
          • Requirements for a Bachelor's Degree
          • Student Status
          • Registration Policies
          • Grading Policies
          • Academic Standing Policies
          • External Credit Policies
          • Study Abroad Policies
          • Graduation Policies
        • Student Life Policies
      • Graduate Policies
        • Graduate-​Specific Policies
        • Academic Policies
        • Admissions and Finances
        • Student Life
          • International Graduate Students
      • Departments, Program Requirements, and Courses
        • Applied Mathematics and Statistics
          • Applied Mathematics and Statistics, Bachelor of Arts
          • Applied Mathematics and Statistics, Bachelor of Science
          • Applied Mathematics and Statistics, Master of Science in Engineering
          • Applied Mathematics and Statistics, Minor
          • Applied Mathematics and Statistics, PhD
          • Data Science, Master's Degree
          • Financial Mathematics, Master of Science in Engineering
        • Biomedical Engineering
          • Bioengineering Innovation and Design, Master of Science in Engineering
          • Biomedical Engineering, Bachelor of Arts
          • Biomedical Engineering, Bachelor of Science
          • Biomedical Engineering, Master of Science in Engineering
          • Biomedical Engineering, PhD through the School of Medicine
        • Center for Leadership Education
          • Accounting and Financial Management, Minor
          • Engineering Management, Master of Science
          • Entrepreneurship and Management, Minor
          • Leadership Studies, Minor
          • Marketing and Communications, Minor
          • Professional Communication Program
          • Professional Development Program
        • Chemical and Biomolecular Engineering
          • Chemical and Biomolecular Engineering, Bachelor of Science
          • Chemical and Biomolecular Engineering, Master of Science in Engineering
          • Chemical and Biomolecular Engineering, PhD
        • Civil &​ Systems Engineering
          • Civil Engineering, Bachelor of Science
          • Systems Engineering, Bachelor of Science
          • Civil Engineering, Master of Science in Engineering (MSE)
          • Civil Engineering, Minor
          • Civil and Systems Engineering, PhD
          • Systems Engineering, Master of Science in Engineering (MSE)
        • Computational Medicine
          • Computational Medicine, Minor
          • Computational Medicine, Pre-​Doctoral Training Program
        • Computer Science
          • Computer Science, Bachelor of Arts
          • Computer Science, Bachelor of Science
          • Computer Science, Master of Science in Engineering
          • Computer Science, Minor
          • Computer Science, PhD
        • Doctor of Engineering
          • Engineering, Doctor of Engineering
        • Electrical and Computer Engineering
          • Computer Engineering, Bachelor of Science
          • Electrical and Computer Engineering, Master of Science in Engineering
          • Electrical and Computer Engineering, PhD
          • Electrical Engineering, Bachelor of Arts
          • Electrical Engineering, Bachelor of Science
          • Energy, Minor
        • Environmental Health and Engineering
          • Engineering for Sustainable Development, Minor
          • Environmental Engineering, Bachelor of Science
          • Environmental Engineering, Minor
          • Environmental Sciences, Minor
          • Geography and Environmental Engineering, Master of Arts
          • Geography and Environmental Engineering, Master of Science
          • Geography and Environmental Engineering, Master of Science in Engineering
          • Geography and Environmental Engineering, PhD
          • Occupational and Environmental Hygiene, Master of Science
        • General Engineering
          • General Engineering, Bachelor of Arts
        • Information Security Institute
          • Security Informatics, Master of Science
          • Security Informatics, Master of Science/​Applied Mathematics and Statistics, Master of Science in Engineering Dual Master's Program
          • Security Informatics, Master of Science/​Computer Science, Master of Science in Engineering Dual Master's Program
        • Materials Science and Engineering
          • Materials Science and Engineering, Bachelor of Science
          • Materials Science and Engineering, Master of Science in Engineering
          • Materials Science and Engineering, PhD
        • Mechanical Engineering
          • Engineering Mechanics, Bachelor of Science
          • Mechanical Engineering, Bachelor of Science
          • Mechanical Engineering, Master of Science in Engineering
          • Mechanical Engineering, PhD
        • NanoBioTechnology
          • Nano-​Biotechnology, Certificate of Advanced Study
        • Robotics and Computational Sensing
          • Computer Integrated Surgery, Minor
          • Robotics, Master of Science in Engineering
          • Robotics, Minor
      • Multi-​School Programs of Study
        • Business, Minor
        • Peabody-​Homewood Double Degree Program
        • Space Science and Engineering
    • Part-​Time, Online Graduate Programs (Engineering for Professionals)
      • Academic Policies
        • Academic Calendar
        • Academic Regulations
        • Registration Policies
        • Tuition and Fees
      • Admission Requirements
      • Applied and Computational Mathematics
        • Applied and Computational Mathematics, Master of Science
        • Applied and Computational Mathematics, Post-​Master’s Certificate
      • Applied Biomedical Engineering
        • Applied Biomedical Engineering, Graduate Certificate
        • Applied Biomedical Engineering, Master of Science
        • Applied Biomedical Engineering, Post-​Master’s Certificate
      • Applied Physics
        • Applied Physics, Master of Science
        • Applied Physics, Post-​Master’s Certificate
      • Artificial Intelligence
        • Artificial Intelligence, Graduate Certificate
        • Artificial Intelligence, Master of Science
      • Chemical and Biomolecular Engineering
        • Chemical and Biomolecular Engineering, Master of Chemical and Biomolecular Engineering
      • Civil Engineering
        • Civil Engineering, Graduate Certificate
        • Civil Engineering, Master of Civil Engineering
      • Computer Science
        • Computer Science, Master of Science
        • Computer Science, Post-​Master’s Certificate
      • Cybersecurity
        • Cybersecurity, Master of Science
        • Cybersecurity, Post-​Master’s Certificate
      • Data Science
        • Data Science, Master of Science
        • Data Science, Post-​Master’s Certificate
      • Electrical and Computer Engineering
        • Electrical and Computer Engineering, Graduate Certificate
        • Electrical and Computer Engineering, Master of Science
        • Electrical and Computer Engineering, Post-​Master’s Certificate
      • Engineering Management
        • Engineering Management, Master of Engineering Management
      • Environmental Engineering, Science, and Management Programs
        • Environmental Engineering
          • Environmental Engineering, Graduate Certificate
          • Environmental Engineering, Master of Environmental Engineering
          • Environmental Engineering, Post-​Master’s Certificate
        • Environmental Engineering and Science
          • Environmental Engineering and Science, Graduate Certificate
          • Environmental Engineering and Science, Master of Science
          • Environmental Engineering and Science, Post-​Master’s Certificate
        • Environmental Planning and Management
          • Environmental Planning and Management, Graduate Certificate
          • Environmental Planning and Management, Master of Science
          • Environmental Planning and Management, Post-​Master’s Certificate
        • Climate Change, Energy, and Environmental Sustainability, Graduate Certificate
      • Financial Mathematics
        • Financial Mathematics, Master of Science
        • Financial Risk Management, Graduate Certificate
        • Quantitative Portfolio Management, Graduate Certificate
        • Securitization, Graduate Certificate
      • Healthcare Systems Engineering
        • Healthcare Systems Engineering, Master of Science
      • Information Systems Engineering
        • Information Systems Engineering, Graduate Certificate
        • Information Systems Engineering, Master of Science
        • Information Systems Engineering, Post-​Master’s Certificate
      • Materials Science and Engineering
        • Materials Science and Engineering, Master of Science
      • Mechanical Engineering
        • Mechanical Engineering, Master of Science
        • Mechanical Engineering, Post-​Master’s Certificate
      • Occupational and Environmental Hygiene
        • Occupational and Environmental Hygiene, Master of Science
      • Robotics and Autonomous Systems
        • Robotics and Autonomous Systems, Master of Science
      • Space Systems Engineering
        • Space Systems Engineering, Master of Science
      • Systems Engineering
        • Systems Engineering, Graduate Certificate
        • Systems Engineering, Master of Science
        • Systems Engineering, Master of Science in Engineering (ABET-​accredited)
        • Systems Engineering, Post-​Master’s Certificate
      • Technical Management
        • Technical Management, Graduate Certificate
        • Technical Management, Post-​Master’s Certificate
  • Zanvyl Krieger School of Arts and Sciences
    • Full-​time, On-​campus Undergraduate and Graduate Programs (Homewood)
      • Undergraduate Policies
        • Academic Policies
          • Requirements for a Bachelor's Degree
          • Student Status
          • Registration Policies
          • Grading Policies
          • Academic Standing Policies
          • External Credit Policies
          • Study Abroad Policies
          • Graduation Policies
        • Student Life Policies
      • Graduate Policies
        • Academic Policies
        • Admissions and Finances
        • Graduate-​Specific Policies
        • Student Life
          • International Graduate Students
      • Departments, Program Requirements, and Courses
        • Anthropology
          • Anthropology, Bachelor of Arts
          • Anthropology, Minor
          • Anthropology, PhD
        • Archaeology
          • Archaeology, Bachelor of Arts
        • Behavioral Biology Program
          • Behavioral Biology, Bachelor of Arts
        • Bioethics
          • Bioethics, Minor
        • Biology
          • Biology, Bachelor of Arts
          • Biology, Bachelor of Arts/​Master of Science
          • Cellular, Molecular, Developmental Biology and Biophysics, PhD
          • Molecular &​ Cellular Biology, Bachelor of Science/​Master of Science
          • Molecular and Cellular Biology, Bachelor of Science
        • Biophysics
          • Biophysics, Bachelor of Arts
          • Biophysics, Fifth-​Year Master’s Degree
          • Biophysics, PhD -​ Jenkins Biophysics Program
          • Biophysics, PhD -​ Program in Molecular Biophysics
          • Biophysics, PhD -​ The Program in Cell, Molecular Developmental Biology and Biophysics
        • Center for Africana Studies
          • Africana Studies, Bachelor of Arts
          • Africana Studies, Minor
        • Center for Language Education
        • Chemistry
          • Chemistry, Bachelor of Arts
          • Chemistry, PhD
        • Classics
          • Classics, Bachelor of Arts
          • Classics, Bachelor of Arts/​Master of Arts
          • Classics, Minor
          • Classics, PhD
        • Cognitive Science
          • Cognitive Science, Bachelor of Arts
          • Cognitive Science, Master of Arts
          • Cognitive Science, PhD
          • Linguistics, Minor
        • Comparative Thought and Literature
          • Honors Program in the Humanities
          • Humanistic Studies, PhD
        • Earth and Planetary Sciences
          • Earth and Planetary Sciences, PhD
          • Earth and Planetary Sciences, Bachelor of Arts
          • Earth and Planetary Sciences, Minor
          • Energy, Minor
          • Environmental Science, Bachelor of Science
          • Environmental Studies, Bachelor of Arts
          • Environmental Studies, Minor
        • East Asian Studies
          • East Asian Studies, Bachelor of Arts
          • East Asian Studies, Minor
        • Economics
          • Economics, Bachelor of Arts
          • Economics, Minor
          • Economics, PhD
          • Financial Economics, Minor
        • English
          • English, Bachelor of Arts
          • English, Minor
          • English, PhD
        • Film and Media Studies
          • Film and Media Studies, Bachelor of Arts
          • Film and Media Studies, Minor
        • History
          • History, Bachelor of Arts
          • History, Bachelor of Arts/​Master of Arts Four-​Year Program
          • History, Minor
          • History, PhD
        • History of Art
          • History of Art, Bachelor of Arts
          • History of Art, Minor
          • History of Art, PhD
          • History of Art, Bachelor of Arts/​Master of Arts
        • History of Science and Technology
          • History of Science and Technology, PhD
          • History of Science, Medicine and Technology, Minor
          • History of Science, Medicine, and Technology, Bachelor of Arts
        • Interdisciplinary Studies
          • Interdisciplinary Studies, Bachelor of Arts
        • International Studies
          • International Studies, Bachelor of Arts
        • Islamic Studies
          • Islamic Studies, Minor
        • Jewish Studies
          • Jewish Studies, Minor
        • Mathematics
          • Mathematics, Bachelor of Arts
          • Mathematics, Minor
          • Mathematics, Bachelor of Arts/​Master of Arts
          • Mathematics, PhD
        • Medicine, Science, and the Humanities
          • Medicine, Science, and the Humanities, Bachelor of Arts
        • Military Science
        • Modern Languages and Literatures
          • French, Bachelor of Arts
          • French, Minor
          • French, PhD
          • German Bachelor of Arts/​Master of Arts
          • German, Bachelor of Arts
          • German, Minor
          • German, PhD
          • Italian, Bachelor of Arts
          • Italian, Minor
          • Italian, PhD
          • Romance Languages, Bachelor of Arts
          • Spanish, Bachelor of Arts
          • Spanish for the Professions, Minor
          • Spanish Language and Hispanic Cultures, Minor
          • Spanish, PhD
        • Museums and Society
          • Museums and Society, Minor
        • Music
          • Music, Minor
        • Natural Sciences Area Major
          • Natural Sciences Area, Bachelor of Arts
        • Near Eastern Studies
          • Near Eastern Studies, Bachelor of Arts
          • Near Eastern Studies, Minor
          • Near Eastern Studies, PhD
        • Neuroscience
          • Neuroscience, Bachelor of Science
          • Neuroscience, Bachelor of Science/​Master of Science
        • Philosophy
          • Philosophy, Bachelor of Arts
          • Philosophy, Bachelor of Arts/​Master of Arts
          • Philosophy, Minor
          • Philosophy, PhD
        • Physics and Astronomy
          • Astronomy and Astrophysics, PhD
          • Physics, Bachelor of Arts
          • Physics, Bachelor of Science
          • Physics, Bachelor of Science/​Master of Science
          • Physics, Minor
          • Physics, PhD
        • Political Science
          • Political Science, Bachelor of Arts
          • Political Science, PhD
        • Program in Latin American Studies
          • Latin American Studies, Minor
        • Psychological and Brain Sciences
          • Psychology, Bachelor of Arts
          • Psychology, Minor
          • Psychology, PhD
        • Public Health Studies
          • Public Health Studies, Bachelor of Arts
        • Social Policy
          • Social Policy, Minor
        • Sociology
          • Sociology, Bachelor of Arts
          • Sociology, PhD
          • Sociology, PhD/​Applied Mathematics and Statistics, MSE Joint Program
        • Space Science and Engineering
          • Space Science and Engineering, Minor
        • Study of Women, Gender, and Sexuality
          • Women, Gender, and Sexuality, Minor
        • Theatre Arts and Studies
          • Theatre Arts and Studies, Minor
        • Visual Arts
          • Visual Arts, Minor
        • Writing Seminars
          • Writing Seminars Minor
          • Writing Seminars, Bachelor of Arts
          • Writing Seminars, Master of Fine Arts
      • Multi-​School Programs of Study
    • Graduate and Professional Programs (Advanced Academic Programs)
      • About Krieger School of Arts and Sciences
      • Administration and Faculty
      • Enrollment Services
        • Academic Regulations for Online Courses
        • Academic Structure
        • Admission
        • Alumni Benefits
        • Grades /​ Performance /​ Conduct
        • Graduation Requirements
        • Registration
        • Tuition and Fees
      • Programs
        • Applied Economics, Master of Science
          • Applied Economics, MS/​ Investment Certificate
          • Applied Economics, MS/​Financial Management Certificate
        • Applied Economics, MS/​MBA Dual Degree
        • Center for Advanced Governmental Studies
          • Data Analytics and Policy, Master of Science
            • Data Analytics and Policy, MS/​Intelligence, Certificate
          • Data Analytics and Policy, Certificate
          • Geospatial Intelligence, Master of Science
          • Global Security Studies, Master of Arts
            • Global Security Studies, MA/​Intelligence, Certificate
          • Government, MA/​MBA
          • Government, Master of Arts
            • Government, MA/​Intelligence, Certificate
          • Intelligence Analysis, Master of Science
          • Intelligence, Certificate
          • Non-​Governmental Organization (NGO) Management, Master of Arts
          • Non-​Profit Management, Master of Arts
          • Nonprofit Management, Certificate
          • Public Management, Master of Arts
            • Public Management, MA/​Data Analytics and Policy, Certificate
            • Public Management, MA/​Intelligence, Certificate
            • Public Management, MA/​Nonprofit Management, Certificate
        • Center for Biotechnology Education
          • Bioinformatics, Master of Science
          • Biotechnology Education, Certificate
          • Biotechnology Enterprise, Certificate
          • Biotechnology, Master of Science
          • Biotechnology, Master of Science/​MBA
          • Food Safety Regulation, Master of Science
          • Individualized Genomics and Health, Master of Science
          • Master of Biotechnology Enterprise and Entrepreneurship
          • Regulatory Science, Master of Science
          • Sequence Analysis and Genomics, Post-​Master’s Certificate
        • Communication, Master of Arts
        • Communication, Master of Arts/​MBA
          • Communication, Master of Arts/​Nonprofit Management, Certificate
        • Cultural Heritage Management, Master of Arts
          • Cultural Heritage Management, MA/​Digital Curation, Certificate
          • Cultural Heritage Management, MA/​Nonprofit Management, Certificate
        • Digital Curation, Certificate
        • Energy Policy and Climate, Master of Science
        • Environmental Sciences and Policy, Master of Science
          • MS in Environmental Sciences and Policy/​Geographic Information Systems, Certificate
        • Film and Media, Master of Arts
        • Geographic Information Systems, Master of Science
          • Geographic Information Systems, Certificate
        • Master of Liberal Arts
        • Museum Studies, Master of Arts
          • Museum Studies, MA/​Digital Curation, Certificate
          • Museum Studies, MA/​Nonprofit Management, Certificate
        • Organizational Leadership, Master of Science
        • Quantitative Methods in Applied Economics, Post-​Master’s Certificate
        • Regenerative and Stem Cell Technologies, Master of Science
        • Research Administration, Master of Science
        • Science Writing, Master of Arts
          • Science Writing, Certificate
        • Teaching Writing, Master of Arts
          • Teaching Writing, Certificate
        • Writing, Master of Arts
        • Office of Summer and Intersession Programs
  • Course Descriptions
    • AS.010 (History of Art)
    • AS.020 (Biology)
    • AS.030 (Chemistry)
    • AS.040 (Classics)
    • AS.050 (Cognitive Science)
    • AS.060 (English)
    • AS.061 (Film and Media Studies)
    • AS.070 (Anthropology)
    • AS.080 (Neuroscience)
    • AS.100 (History)
    • AS.110 (Mathematics)
    • AS.130 (Near Eastern Studies)
    • AS.136 (Archaeology)
    • AS.140 (History of Science, Medicine, and Technology)
    • AS.145 (Medicine, Science and the Humanities)
    • AS.150 (Philosophy)
    • AS.171 (Physics &​ Astronomy)
    • AS.180 (Economics)
    • AS.190 (Political Science)
    • AS.192 (International Studies)
    • AS.194 (Islamic Studies)
    • AS.196 (Agora Institute)
    • AS.200 (Psychological &​ Brain Sciences)
    • AS.210 (Modern Languages &​ Literatures)
    • AS.220 (Writing Seminars)
    • AS.225 (Theatre Arts &​ Studies)
    • AS.230 (Sociology)
    • AS.250 (Biophysics)
    • AS.270 (Earth &​ Planetary Sciences)
    • AS.280 (Public Health Studies)
    • AS.290 (Behavioral Biology)
    • AS.300 (Comparative Thought and Literature)
    • AS.310 (East Asian Studies)
    • AS.360 (Interdepartmental)
    • AS.361 (Program in Latin American Studies)
    • AS.362 (Center for Africana Studies)
    • AS.363 (Study of Women, Gender, &​ Sexuality)
    • AS.370 (Center for Language Education)
    • AS.371 (Art)
    • AS.374 (Military Science)
    • AS.376 (Music)
    • AS.389 (Program in Museums and Society)
    • AS.410 (Biotechnology)
    • AS.420 ( Environmental Sciences)
    • AS.425 ( Energy Policy and Climate)
    • AS.430 ( Geographic Information Systems)
    • AS.440 ( Applied Economics)
    • AS.450 ( Liberal Arts)
    • AS.455 ( Film and Media)
    • AS.460 ( Museum Studies)
    • AS.465 ( Cultural Heritage Management)
    • AS.470 ( Government)
    • AS.472 ( Geospatial Intelligence)
    • AS.475 ( Research Administration)
    • AS.480 ( Communication)
    • AS.485 ( Organizational Leadership)
    • AS.490 ( Writing)
    • AS.491 ( Science Writing)
    • AS.492 ( Teaching Writing)
    • AS.492 (Non-​Departmental)
    • AS.990 (-​JHU Department)
    • AS.999 (AAP)
    • BU.001 (Graduate Business)
    • BU.001 (MBA)
    • BU.120 (Management)
    • BU.132 (Real Estate)
    • BU.210 (Finance)
    • BU.300 (Information Systems)
    • BU.410 (Marketing)
    • BU.510 (Quantitative Methods)
    • BU.550 (Business of Health)
    • BU.610 (Operations Management)
    • ED. (Education)
    • EN.500 (General Engineering)
    • EN.510 (Materials Science &​ Engineering)
    • EN.515 (Materials Science and Engineering)
    • EN.520 (Electrical &​ Computer Engineering)
    • EN.525 (Electrical and Computer Engineering)
    • EN.530 (Mechanical Engineering)
    • EN.535 (Mechanical Engineering)
    • EN.540 (Chemical &​ Biomolecular Engineering)
    • EN.545 (Chemical and Biomolecular Engineering)
    • EN.553 (Applied Mathematics &​ Statistics)
    • EN.555 (Financial Mathematics)
    • EN.560 (Civil Engineering)
    • EN.565 (Civil Engineering)
    • EN.570 (Environmental Health and Engineering)
    • EN.575 (Environmental Engineering and Science)
    • EN.575 (Environmental Engineering)
    • EN.575 (Environmental Planning and Management)
    • EN.580 (Biomedical Engineering)
    • EN.585 (Applied Biomedical Engineering)
    • EN.595 (Engineering Management)
    • EN.601 (Computer Science)
    • EN.605 (Computer Science)
    • EN.615 (Applied Physics)
    • EN.625 (Applied and Computational Mathematics)
    • EN.635 (Information Systems Engineering)
    • EN.645 (Systems Engineering)
    • EN.650 (Information Security Institute)
    • EN.655 (Healthcare Systems Engineering)
    • EN.660 (Center for Leadership Education)
    • EN.670 (Institute for NanoBio Technology)
    • EN.675 (Space Systems Engineering)
    • EN.685 (Data Science)
    • EN.695 (Cybersecurity)
    • EN.700 (Doctor of Engineering)
    • ME.100 (Biophsyics)
    • ME.110 (Cell Biology)
    • ME.120 (Art as Applied to Medicine)
    • ME.130 (Functional Anatomy and Evolution)
    • ME.140 (Gynecology and Obstetrics)
    • ME.150 (History of Medicine)
    • ME.200 (Neurology)
    • ME.210 (Biomedical Engineering)
    • ME.220 (Dermatology)
    • ME.250 (Medicine)
    • ME.250 (Health Sciences Informatics)
    • ME.260 (Molecular Biology and Genetics)
    • ME.280 (Ophthalmology)
    • ME.290 (Otolaryngology)
    • ME.300 (Pathology)
    • ME.320 (Pediatrics)
    • ME.330 (Pharmacology and Molecular Sciences)
    • ME.340 (Biological Chemistry)
    • ME.360 (Physiology)
    • ME.370 (Psychiatry)
    • ME.380 (Surgery)
    • ME.381 (Plastic Surgery)
    • ME.390 (Neurosurgery)
    • ME.400 (Orthopedic Surgery)
    • ME.420 (Radiology)
    • ME.440 (Neuroscience)
    • ME.520 (Emergency Medicine)
    • ME.510 (Oncology)
    • ME.560 (Urology)
    • ME.570 (Anesthesiology)
    • ME.580 (Biomedical Engineering)
    • ME.600 (Health Sciences Informatics)
    • ME.680 (Comparative Medicine)
    • ME.700 (Immunology)
    • ME.710 (Human Genetics)
    • ME.711 (Berman Bioethics Institute)
    • ME.712 (The Welch Center)
    • ME.714 (The Bloomberg School of Public Health)
    • ME.715 (Non-​Departmental)
    • ME.716 (Physical Medicine &​ Rehabilitation)
    • ME.717 (Radiation Oncology)
    • ME.718 (Institute of Genetic Medicine)
    • ME.800 (Interdepartmental)
    • NR (Nursing)
    • PH.120 (Biochemistry and Molecular Biology)
    • PH.140 (Biostatistics)
    • PH.220 (International Health)
    • PH.260 (Molecular Microbiology and Immunology)
    • PH.300 (Health Policy and Management)
    • PH.330 (Mental Health)
    • PH.340 (Epidemiology)
    • PH.380 (Population Family and Reproductive Health)
    • PH.390 (Clinical Investigation)
    • PH.410 (Health Behavior and Society)
    • PH.550 (Extradepartmental Studies)
    • PH.600 (Online Programs for Applied Learning)
    • PH.700 (Berman Institute)
    • PY.010 (Studio Lessons)
    • PY.113 (Recitals)
    • PY.123 (General Studies)
    • PY.123 (Professional Studies)
    • PY.250 (Humanities -​ Language)
    • PY.260 (Humanities -​ Liberal Arts)
    • PY.310 (Composition)
    • PY.320 (New Media)
    • PY.330 (Conducting)
    • PY.350 (Computer Music)
    • PY.380 (Historical Performance)
    • PY.410 (Brass)
    • PY.415 (Percussion)
    • PY.420 (Harp)
    • PY.425 (Strings)
    • PY.430 (Woodwinds)
    • PY.450 (Ensemble Arts)
    • PY.450 (Piano/​Keyboard)
    • PY.460 (Organ)
    • PY.470 (Guitar)
    • PY.510 (Music Education)
    • PY.520 (Pedagogy)
    • PY.530 (Voice)
    • PY.540 (Opera)
    • PY.550 (Recording Arts and Sciences)
    • PY.570 (Jazz)
    • PY.610 (Musicology)
    • PY.710 (Music Theory)
    • PY.715 (Music Theory -​ ET/​SS)
    • PY.715 (Music Theory -​ Keyboard Studies)
    • PY.800 (Dance)
    • PY.910 (Ensembles -​ Large)
    • PY.950 (Ensembles -​ Small/​Chamber)
    • SA.100 (Core Courses)
    • SA.310 (International Economics)
    • SA.500 (Development, Climate &​ Sustainability)
    • SA.501 (Technology &​ Culture)
    • SA.503 (States Markets Institutions)
    • SA.510 (International Economics, &​ Finance)
    • SA.550 (Africa)
    • SA.551 (The Americas)
    • SA.552 (Asia)
    • SA.553 (China)
    • SA.554 (Europe &​ Eurasia)
    • SA.555 (The Middle East)
    • SA.556 (The United States)
    • SA.600 (International Relations)
    • SA.620 (Global Policy)
    • SA.630 (Global Risk)
    • SA.502 (Security Strategy, &​Statecraft)
  • Course Search
    • /​course-​search/​api/​
  • Catalogue Contents
  • Catalogue Archives
  • Amendments
  • Home›
  • School of Medicine›
  • Graduate Programs›
  • Immunology, PhD
  • Overview
  • Admission
  • Requirements

Department website: https://gradimmunology.med.som.jhmi.edu/

For more complete information and to apply to the Graduate Program in Immunology see the department website.

The Graduate Program in Immunology began in 1982 with two students and a handful of faculty. Since then we have grown to 36 students currently in training and 46 faculty. The faculty research interests span the entire gamut of Immunology, from basic mechanisms such as somatic hypermutation and gene rearrangement, molecular mechanisms of antigen processing and presentation, molecular signals for immune cell development and differentiation to translational research in the fields of cancer vaccines, allergy, infectious disease, immune engineering, autoimmunity and transplantation.

We are fortunate to have a number of centers of excellence in Basic Immunology and Cellular Engineering, Microbial Immunology, Autoimmunity and Cancer Immunology. This, combined with world-class facilities in genomics, genetics, proteomics, cell imaging and bioinformatics means almost limitless opportunities for students to do cutting edge and creative Immunology research. It is indeed an exciting time for Immunology at Hopkins.

 We are able to offer one of the most rigorous and expansive scientific training arenas in the world. The large number of accomplished biomedical scientists, the focus on interdisciplinary training, the availability of state of the art research facilities and a collegial environment gives our students a unique opportunity to develop as the next generation of leaders in Immunology. Recent program graduates have become post-doctoral research fellows in major labs all over the world. Many of our alumni have gone on to significant positions as successful independent researchers at universities, research institutes, government laboratories and biotechnology enterprises. Also, a number have distinguished themselves in public policy, intellectual property and scientific writing. All of our alumni have gone on to fulfilling careers in the biomedical field and for that we are grateful and very proud.

Fellowships

Financial Support:

Students accepted into the program will have their tuition, fees, and medical insurance (including dental and vision) paid. Students will receive a competitve stipend to cover living expenses. Support for the Graduate Program in Immunology is derived from government and private sources and individual research grants.

Note to International Applicants:

We are very much interested in attracting talented students from other countries to our program. However, there are some practical issues that you should be aware of before applying. We have a limited ability to support students who are not citizens or permanent residents of the United States. Our training program is funded by a training grant from the United States government. This grant will only support US citizens or permanent residents. Only very rarely do we have funds that can be used to support citizens of other countries.

We highly recommend that foreign applicants seek other possible means of supporting the cost of graduate study. Support could come from: (a) a scholarship from your government or (b) from a "special" foundation award. In your application, please discuss the possibility of obtaining support from one of these sources. Final admission to the program requires documentation that the appropriate financial resources are available. We highly recommend that you review the NAFSA: Association of International Educators website at www.nafsa.org. NAFSA does not offer financial assistance, but will offer some suggestions to help your search for financial aid for study in the United States.

Acceptance into the Graduate Program in Immunology is dependent on an invited personal interview. Foreign applicants, on invitation, should be prepared to travel to Baltimore, Maryland at their own expense. Once in Baltimore, the program will pay for local transportation, meals and accommodations.

Admission Requirements

Undergraduate Degree 

Candidates for admission should hold a bachelor’s degree or higher with training in one of the following, or related, areas:
General Biology
Biochemistry
Physics
Calculus
Organic/Inorganic Chemistry
Courses in Immunology, Biochemistry & Molecular Biology are recommended, but not required.

Transcripts

Transcripts of all college and university study (undergraduate and graduate) are required and must be uploaded to the application. If you have attended more than one institution, transcripts from each institution must be received for your application to be considered complete. International transcripts must be officially translated into English. If you have not yet completed your Bachelor’s degree, upload a copy of your current transcript, showing in-progress courses. Official transcripts for in-progress courses can be uploaded to the online application in the Transcript Upload section when they become available, after you submit. Applicants should also upload a list of any current courses, and courses that will be taken before beginning graduate study that do not appear on their transcripts. Do not mail documents to the Office of Graduate Student Affairs unless requested or admitted.
If admitted, an official transcript from each institution you have attended showing proof of graduation and degree conferral will be required prior to matriculation. To be considered official, final transcripts must be sent: 1) by mail, directly from the institution in a sealed envelope: 

Office of Graduate Biomedical Education
1830 E. Monument Street, Suite 620
Baltimore, MD 21287

If your institution is sending an E-transcript, please make sure they are sending the notifications to:  Gradadmissions@jhmi.edu 

Letters of Recommendation 

Three letters of recommendation must be submitted through the online recommendation system. Applicants will need to obtain the name and email address of the recommenders when completing the application. Please request your letters of recommendation from faculty members or other professionals who are acquainted with you and your academic work. These letters should comment on your aptitude and promise for independent research.

GRE (no longer required for admission)

As of October 2019, The Graduate Program in Immunology will no longer require the Graduate Record Exam (GRE) for admission. 

TOEFL Scores (Test of English as a Foreign Language) and IELTS (International English Language Testing System)

All levels of graduate education at Johns Hopkins require proficiency in the English language. It is highly recommended that applicants for whom English is a second language take the TOEFL/IELTS to demonstrate this proficiency. In general, foreign students admitted to the Graduate Program in Immunology have achieved TOEFL scores over 550.
You are exempt from having to take the TOEFL/IELTS if you hold a degree from a U.S. University/College or have attended a minimum of 2 years at an English speaking University/College. TOEFL/IELTS scores are valid for 2 years from expected entrance date.
Please use the Johns Hopkins School of Medicine code (5316) when submitting scores.

Application Fee

Application fee is $115.

Criminal Background Check

All Johns Hopkins University graduate students who are accepted will be subject to criminal background investigations. Generally, all offers of admission to School of Medicine degree programs will be conditioned on satisfactory criminal background investigations.

Program Requirements 

In the first year of study, each student takes a core set of courses emphasizing basic molecular principles and how they apply to understanding immune function.

Required Coursework

Course List
Code Title Credits
ME.100.709Macromolecular Structure and Analysis1.5
ME.100.710Biochemical and Biophysical Principles1.5
ME.110.728Cell Structure and Dynamics1.5
ME.250.703Graduate Immunology4
ME.250.709Immunology Course Discussion1
ME.250.722Autoimmunity1
ME.250.804Introduction to Immunology Research (Parts I and II)1
ME.260.709Molecular Biology and Genomics1.5
ME.260.708Fundamentals of Genetics2
ME.260.802Special Studies and Research0
ME.360.728Pathways and Regulation2
ME.800.707Computational Biology and Bioinformatics0.5

Laboratory Rotations

During the first year, each student engages in three short-term research projects. Each project lasts approximately three months and is carried out under the direction of a faculty member. Additional rotations may be scheduled based on student interest. The projects are designed to give you an introduction to experimental research and an opportunity to learn more about specific areas of immunology prior to choosing a thesis laboratory and project. After completing the research rotations, the student will select a mentor for their thesis project.

2nd Year

Elective Courses

Students are required to take five elective courses prior to graduation. Students can begin taking courses in the spring of their first year, but most students opt to start in their second year. There are many available advanced level graduate courses offered in the School of Medicine, Bloomberg School of Public Health, and the Kreiger School of Arts and Sciences. The Immunology Program specifically offers several courses, including Tumor Immunology and Immunotherapy, Immunometabolism, Translational Immunology, HIV Biology and more. Many of these courses utilize small group discussions, in which students read and discuss current and seminal research papers on the selected topic.

Oral Examination

In the fall of the second year of study, trainees will take the Graduate Board Oral Examination. By the time the students take this exam, they will have successfully completed all required coursework. This examination serves as a means of evaluating the student’s cumulative knowledge in biochemistry, cellular and molecular biology, biophysics, genetics, and immunology and their preparedness to carry out research for the Ph.D. degree. The exam may cover the student's proposed dissertation topic, but this is not the focus of the exam.

Formation of Thesis Committee 

In the spring of the second year of study, students will bring together 4-5  faculty members, including their mentor, to serve as their thesis advisory committee. The purpose of the thesis committee is to help the student move their research forward, provide networking opportunities and career development advice and to ensure the student successfully completes their degree.  At the initial meeting, the student will prepare a written research proposal in the form of an NIH grant.

3rd Year and Beyond

Elective Courses

Students are required to take five elective courses prior to graduation. Students can begin taking courses in the spring of their first year, but most students opt to start in their second year. There are many available advanced level graduate courses offered in the School of Medicine, Bloomberg School of Public Health, and the Kreiger School of Arts and Sciences. The Immunology Program specifically offers several courses, including Tumor Immunology and Immunotherapy, Immunometabolism, Translational Immunology, HIV Biology and more. Many of these courses utilize small group discussions, in which students read and discuss current and seminal research papers on the selected topic. 

Thesis Meetings

Students are required to have at least ONE thesis meeting per year, but are welcome to have more if desired. At each committee meeting, the student should present their research work, roughly following the format of the written proposal. The discussion can be, and often is, open-ended in nature. The student should be prepared to discuss:
1. Background and significance
2. Specific goals of the research (specific aims) 
3. Work accomplished to date, including pertinent experiments that "did not work"
4. Future experiments (long and short term) 

Thesis and Final Seminar 

Upon completion of the thesis research, each student must prepare a formal written thesis, based on the guidelines provided by the Graduate Board of the University. Two readers must find the written thesis acceptable: the thesis advisor and another member of the Thesis Advisory Committee. Students must also present a formal public seminar on their research. The program office will schedule the final seminar. All University guidelines for thesis preparation must be met. More detailed information on this process is available in the program office.

Required Program Events

The Graduate Program in Immunology offers a wide range of activities that serve to enrich the training experience. All students are required to participate in the program activities throughout their graduate career and this is a vital aspect of the training program. 

Journal Club 

Immunology Journal Club is intended to provide all graduate students in the Graduate Immunology Program the habit of reading a diverse range of immunology journal articles early in their graduate careers. Presenters are generally encouraged to present new developments and findings that are less related to their research focus. This will allow participants to explore new areas of immunology, familiarize themselves with key investigators in the immunology field, and to develop sharp and valid criticism of sound experiments. This is an invaluable opportunity to keep abreast of new advances as well as hone one's presentation skills in an informal setting.

Annual Immunology Retreat

The Graduate Program in Immunology holds an annual fall retreat. This is a one-day activity at an off-site location. The day includes poster presentations by all Immunology trainees (3rd year and above), a series of 5-6 short student oral presentations (4th years), and a keynote address by a distinguished invited speaker. This meeting provides students with an excellent opportunity to hone presentation skills in an informal retreat setting.

Immunology Floor Meeting 

Students and faculty of the Graduate Program in Immunology gather regularly on Thursdays for the “Immunology Floor Meeting.” At these meetings, two speakers from different labs will each give a 30 minute presentation on their research in progress. These presentations are designed to allow for the exchange of ideas in an informal atmosphere. The idea is to have people present, as if they were presenting at lab meeting, so we can exchange ideas at the frontier of our research. These are not to be highly polished presentations but a description of your project as it stands, with people encouraged to talk not just about exciting new results, but also about difficulties, troubleshooting, new techniques being attempted, ideas, etc. The speakers can be graduate students, post-docs, or even faculty members, and we hope the venue will be an excellent opportunity for honing our speaking skills.

Immunology Forum 

Students are required to attend the Johns Hopkins Immunology Forum that is scheduled regularly on Tuesdays at 4PM. As an added bonus for the students, we provide lunch for the speaker and the students on Forum days. Seminar speakers come from various institutions in the United States and abroad and meet with faculty and trainees. Immunology students are given the opportunity to invite several speakers each year. On this date, the students extend the invitation, organize the schedule, and take them to dinner. The seminar coordinator will assist in this. Collectively these activities provide a powerful training opportunity, allowing trainees to not only hear an interesting seminar, but also to develop the skills and confidence enabling them to enter into scientific dialogue with a gifted scientist.

Johns Hopkins University
  • Johns Hopkins University
  • Baltimore, MD
  • 410-516-8000
  • © 2019 Johns Hopkins University. All rights reserved.
  • About Us
  • Academics
  • Schools & Divisions
  • Admissions & Aid
  • Research & Faculty
  • Campus Life
Back to top

Print Options

  • Send Page to Printer

    Print this page.

  • Download Page (PDF)

    The PDF will include all information unique to this page.

  • Download PDF of the entire Catalogue

    All pages in the catalogue.